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ProxiesHoneybeeCockroachFruit FlyAntsC elegansCrabCrayfishEarthwormSea hareMoon jellyfishSpidersOctopusesPlants ProkaryotProtistChickenCowHumanSometimes operates unconscious in HU/ ANSoldierflyCyprinidaeSalmonidaeShrimpSilk wormPig
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Noxious Stimuli Reactions
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Physiological responses to nociception or handlingLikely YesUnknownLikely YesUnknownLikely YesLikely yesLikely YesLean YesLean YesUnknownUnknownLikely YesLikely YesUnknownLean YesLikely YesLikely YesYesLikely YesUnknownLikely YesLikely yesLean YesUnknownLikely Yes
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Protective behaviorLean YesLean YesLean YesLean YesUnknownLikely yesLean YesUnknownUnknownUnknownUnknownLikely YesLikely NoUnknownUnknownLikely YesLikely YesYesLikely YesUnknownLikely YesLikely yesLean YesLean YesLikely Yes
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Defensive behavior/fighting backLikely YesLikely YesLikely YesLikely YesUnknownLikely yesLikely YesUnknownLikely YesLikely YesLikely YesLikely YesLean NoUnknownUnknownLikely YesLikely YesYesLean NoUnknownLikely YesLikely yesUnknownLean yesLikely Yes
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Noxious stimuli related vocalizationsLikely YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely NoUnknownUnknownLikely NoLikely NoLikely NoLikely YesLikely YesYesLikely YesUnknownLikely YesUnknownUnknownLean yesLikely Yes
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Movement away from noxious stimuliLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLikely YesLikely YesLikely YesLikely YesLikely NoLikely YesLikely YesLikely YesLikely YesYesLikely YesUnknownLikely YesLikely yesLikely YesLean yesLikely Yes
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Motivational Tradeoffs
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Paying a cost to receive a rewardLean YesUnknownLikely YesLean YesUnknownLean yesLean YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownLean YesLikely YesLean YesYesLikely NoUnknownLikely YesLikely YesUnknown UnknownLikely Yes
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Paying a cost to avoid a noxious stimulusUnknownUnknownUnknownUnknownUnknownLikely yesUnknownUnknownUnknownUnknownUnknownLean YesUnknownUnknownUnknownUnknownLean YesYesLean YesUnknownLean YesLean YesUnknownLean yesLikely Yes
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Self-controlLean YesUnknownUnknownUnknownUnknownLean yesUnknownUnknownUnknownLean NoUnknownLean YesLikely NoLikely NoUnknownLean NoUnknownYesLean YesUnknownLean YesUnknownUnknownUnknownLikely Yes
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Predator avoidance tradeoffsLikely YesUnknownLikely YesUnknownLean yesLikely yesLikely YesLean YesUnknownUnknownUnknownLean YesUnknownUnknownLean NoLikely YesLikely NoYesLean YesUnknownLean YesLean YesLean yesUnknownUnknown
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Selective attention to noxious stimuli over other concurrent eventsUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLean YesUnknownUnknownLikely YesUnknownUnknownUnknownUnknownLikely YesYesLikely NoUnknownUnknownLikely YesLikely yesLean yesUnknown
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Learning Indicators
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Classical conditioningLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLikely YesUnknownLikely YesLikely YesLean YesLean NoLean YesLikely YesLikely YesYesLikely YesLikely YesLikely YesLikely YesLean yesLikely yesLikely Yes
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Operant conditioningLikely YesLikely YesLikely YesLikely YesUnknownLikely yesLikely YesLean YesLikely YesUnknownLikely YesLikely YesUnknownUnknownLean NoLikely YesLikely YesYesLikely YesUnknownLean YesLikely YesUnknownUnknownLikely Yes
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Operant conditioning with unfamiliar actionLikely YesLikely YesLikely YesUnknownUnknownLikely yesLikely YesLean NoUnknownUnknownUnknownLikely YesUnknownUnknownUnknownLikely YesLikely YesYesLean NoUnknownLean YesLikely YesUnknownUnknownLikely Yes
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SensitizationLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLikely YesUnknownLean YesLikely YesLean NoLikely YesLikely YesLikely YesLikely YesYesLikely YesUnknownLean YesUnknownUnknownUnknownLikely Yes
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HabituationLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLikely YesLikely YesLikely YesLikely YesLean YesLean YesLikely YesLikely YesLikely YesYesLikely YesUnknownLikely YesLikely YesUnknownUnknownLikely Yes
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Contextual learningLikely YesLikely YesLikely YesLikely YesUnknownLikely yesLean YesUnknownLean NoUnknownUnknownLean YesUnknownUnknownUnknownUnknownUnknownYesUnknownUnknownUnknownUnknownUnknownUnknownLikely Yes
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Observational or social learningLikely YesLean YesLean YesLean YesLikely NoUnknownLean YesUnknownUnknownLikely NoLean YesLean YesUnknownUnknownUnknownLikely YesLean YesYesLikely YesUnknownLikely YesLikely YesUnknownUnknownLikely Yes
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Taste aversion behaviorLikely YesUnknownLean NoUnknownLean YesLikely yesLikely YesUnknownUnknownUnknownUnknownLean YesLikely NoLikely NoLikely NoLikely YesLikely YesYesLikely YesLean yesLikely YesLikely YesLikely YesLean NoLikely Yes
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Pain relief learningLean yesUnknownLikely YesUnknownUnknownLean yesLean YesUnknownUnknownUnknownUnknownLikely YesUnknownUnknownUnknownUnknownUnknownYesLikely YesUnknownLean YesUnknownLean YesUnknownLikely Yes
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Long-term behavior alteration to avoid noxious stimulus (24+ hours)UnknownLikely YesLikely YesUnknownLikely YesLikely yesLikely YesLean YesLikely YesUnknownLean YesLikely YesUnknownLikely NoLikely NoLikely YesLikely YesYesLean YesUnknownLean YesLean YesUnknownUnknownLikely yes
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Long-term behavior alteration to avoid noxious stimuli (30+ days)UnknownUnknownUnknownUnknownUnknownUnknownUnknownLean YesLean NoUnknownUnknownUnknownUnknownLikely NoLikely NoLikely YesLean YesYesLean YesUnknownUnknownUnknownUnknownUnknownLikely yes
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Cognitive Sophistication
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Uncertainty monitoringLean YesUnknownUnknownLean YesUnknownUnknownLean YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownYesLikely NoUnknownUnknownUnknownLean YesUnknownUnknown
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Self-recognition (mirror test)UnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLean NoLikely NoUnknownUnknownLikely NoLean yesYesLean NoUnknownUnknownUnknownUnknownUnknownLean Yes
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DeceptionUnknownUnknownUnknownLean YesUnknownUnknownLean YesUnknownUnknownUnknownUnknownLean YesLikely NoUnknownUnknownLean YesUnknownYesLikely YesUnknownUnknownUnknown UnknownUnknownLean Yes
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Tool useLean YesUnknownUnknownLikely YesUnknownLean yesUnknownUnknownUnknownUnknownLean NoLikely YesUnknownUnknownUnknownUnknownUnknownYesLean YesUnknownUnknownUnknownUnknownUnknownLean Yes
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Flexible tool useUnknownUnknownUnknownLean YesUnknownLean noUnknownUnknownUnknownUnknownUnknownLean YesUnknownUnknownUnknownUnknownUnknownYesLean NoUnknownUnknownUnknownUnknownUnknownUnknown
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Drug Responses
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Affected by analgesics in a manner similar to humansLikely YesLikely YesLikely YesUnknownLean YesLean yesLikely YesLikely YesLean YesUnknownUnknownLikely YesUnknownUnknownUnknownLikely YesLikely YesYesn/aUnknownLikely YesLikely YesLikely yesUnknownLikely Yes
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Self-administers analgesicsLikely NoUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesUnknownUnknownUnknownLean YesUnknownYesLean NoUnknownLean YesLean YesUnknownUnknownLean Yes
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Affected by recreational drugs in a similar manner to humansLikely YesUnknownLikely YesUnknownLikely YesUnknownLikely YesLean YesUnknownUnknownLean YesLean YesUnknownUnknownLean YesLean NoLikely YesYesn/aUnknownLean YesLikely YesUnknownUnknownLean Yes
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Self-administers recreational drugsLean YesUnknownLikely YesLean YesLikely YesUnknownLean YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLean NoLikely YesYesLean NoUnknownLean YesLean YesUnknownUnknownUnknown
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Affected by antidepressants or anxiolytics in a similar manner to humansUnknownUnknownLikely YesUnknownLean YesLean yesLikely YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesLikely YesYesn/aUnknownLean YesLean YesUnknownUnknownLikely yes
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Navigational Skills
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Navigate known paths/areasLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLean YesLean NoLikely YesLikely YesLikely NoLikely NoLikely YesLikely YesLikely YesYesLean YesLikely YesLikely YesLikely YesLean YesLikely YesLikely Yes
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Navigate unknown paths/areasLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLean YesLean NoLikely YesLikely YesLikely NoLikely NoLikely YesLikely YesLikely YesYesLean YesLikely YesLikely YesLikely YesLean YesLikely YesLikely Yes
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Spatial memoryLikely YesLikely YesLikely YesLikely YesUnknownLikely yesLikely YesUnknownUnknownLikely NoLikely YesLikely YesLikely NoLikely NoLikely YesLikely YesLikely YesYesUnknownUnknownLikely YesLikely YesUnknownUnknownLean Yes
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Detour behaviorLikely YesUnknownLean YesLikely YesUnknownLean yesUnknownUnknownUnknownUnknownLikely YesLikely YesLikely NoLikely NoUnknownLikely YesLikely YesYesLean YesUnknownUnknownUnknownUnknownUnknownLikely Yes
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Mood State Behaviors
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Anhedonia behaviorUnknownUnknownLikely YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesYesLean NoUnknownLean yes UnknownUnknownUnknownLikely Yes
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Learned helplessness behaviorLean YesLikely YesLikely YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesLikely YesYesLean NoUnknownLean yesLean YesUnknownUnknownLikely Yes
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Displacement behaviorLean YesUnknownLean YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesLean YesYesLikely YesUnknownUnknownUnknownLean yesUnknownLikely Yes
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Adverse effects of social isolationLean YesLean NoLean YesLikely YesLikely YesUnknownLikely YesUnknownLean YesUnknownLean YesLean NoUnknownUnknownUnknownLikely YesLikely YesYesLikely YesUnknownLean yesLean YesUnknownLean yesLikely Yes
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Stereotypic behaviorUnknownUnknownUnknownUnknownUnknownUnknownLikely YesUnknownUnknownUnknownUnknownLean YesLikely NoUnknownUnknownLikely YesLikely YesYesLikely YesUnknownUnknownUnknownUnknownUnknownLikely Yes
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Fear/anxiety behaviorLean YesUnknownLikely YesUnknownLean NoLean yesLikely YesUnknownLean YesUnknownUnknownLikely YesUnknownUnknownUnknownLikely YesLikely YesYesLikely YesLean YesLikely yesLikely yesLean YesUnknownLikely Yes
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Play behaviorLean YesUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownUnknownLikely YesUnknownUnknownUnknownLikely YesLikely YesYesLean YesUnknownLean yesLean yesUnknownUnknownLikely Yes
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Anatomical and Evolutionary Features
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Neuron count0.02-1.1 mio~1000000 (brain neurons)~100,000 - 135,000 (brain neurons)Unknown302 (total CNS)0.1 mio0.08 mio~10,000 (brain neurons)10,000 - 20,000 (total CNS)Unknown
~30,000
genus Cupiennius: ~100,000 (total CNS)
~500 mio000220 mio3 billion86.1 ± 8.1 billion (brain neurons)n/a0.3-0.35 mio8-13 mio8-13 mio0.08 mio0.86 mi320-430 mio
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Brain size0.60 - 1.86 mm3.7 to 2 mm wide590 µm × 340 µm × 120 µmAnt brain sizes range from 0.06 mg in Monomorium floricola to 1.730 mg in Paraponera clavata5000µm3Scylla serrata, brain diameters: 1.55mm x 2.08mm

Leptograpsus variegatus, brain diameters: 1.25mm x 1.53mm
UnknownUnknownUnknown00.1-5% of body weightVaries0002.819 g ± 0.200 g~400 g974.9cm^3 - 1498.5cm^3n/aUnknown230623-384614 mm3 0.10 ± 0.04 (olfactory bulb), 0.37 ± 0.16 (telencephalon), 0.78 ± 0.23 (optic tectum), 0.72 ± 0.30 (cerebellum), 0.17 ± 0.07. (hypothalamus)UnknownUnknown95077 mm3
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NociceptionLikely YesLikely YesLikely YesLikely YesLikely YesLikely YesLikely yesLikely YesLikely YesLikely YesLikely YesLikely YesLean YesLikely YesLean YesLikely YesLikely YesYesn/aLean yesLikely yesLikely yesLikely yesLean yesLikely Yes
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Nociception (strict definition)UnknownUnknownLikely YesUnknownLikely YesUnknownLean yesUnknownLikely YesUnknownUnknownLikely YesLikely NoLikely NoLikely NoLikely YesLikely YesYesn/aUnknownLikely yesLikely yesUnknownUnknownLikely Yes
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Opioid-like receptorsLikely YesLikely YesLean YesLean YesLikely YesLikely YesLikely yesLikely YesLikely YesUnknownUnknownLean YesLikely NoLikely NoLikely YesLikely YesLikely YesYesn/aLean yesLikely yesLikely yesLean yesUnknownLikely Yes
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Centralized information processingLikely YesLikely YesLikely YesLikely YesLean NoLikely YesLikely yesLean YesLean YesLean NoLikely YesLikely YesLikely NoLikely NoLikely NoLikely YesLikely YesYesn/aLean yesLikely YesLikely YesLean yesLean yesLikely Yes
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Vertebrate midbrain-like functionLikely YesLikely YesLikely YesLikely YesLikely NoLikely YesLikely yesLean NoUnknownLikely NoLean YesLean YesLikely NoLikely NoLikely NoLikely YesLikely YesYesn/aUnknownLikely YesLikely YesLean NoLean NoLikely Yes
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Last common ancestor with humans (millions of years)79779779779779779779779779782479779714964290148031296n/an/a68043143168068094
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Average Lifespan3w-6m245 to 1878d25-30 daysant queens 10 +/- 6.6y5d2.5-3.5y3-6y1-2 years1y4-8m595d-decades1-2y1d-4500y20min-1h12h-3d7-8 years12-22 years72 yearsn/a4517-20 years , maximum lifespan 47 years in captivity4-5 years average, max. 13 years1-2y34-49d15-20y
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MotileYesYesYesYesYesYesYesYesYesLikely YesYesYesLikely NoLikely YesLikely YesYesYesYesLikely YesYesYesYesYesYesYes
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Distinct sleep-wake statesLikely YesLikely YesLikely YesLikely YesLean YesLean YesLean yesLean YesLikely YesLean YesUnknownLikely YesLikely NoLikely NoLikely NoLikely YesLikely YesYesLikely NoUnknownLikely yesLikely yesUnknownUnknownLikely yes
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