2019-2020 Haplogroup E Tree
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ABCDEFGHIJKLMNOPQRSTUVWXYZAAABACADAEAFAG
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Y-DNA Haplogroup E and its Subclades - 2019-2020
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Downloading this spreadsheet as Excel retains format. Links may not copy to new sheet.
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The entire work is identified by the Version Number and date given on
Main Page.
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Directions for citing the document are listed:
Main Page.
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Version History
Last revision date for this specific page: 1 October 2019
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Because of continuing research, structure of the Y-DNA Haplogroup Tree changes, and will be updated when possible
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LINKS:
Main Page.
Listing Criteria
Papers/Presentations Cited
Y-DNA Tree Trunk
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Glossary
SNP Index
Copyright 2020, International Society of Genetic Genealogy.
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FONT COLORS:
Newly confirmed in 2019 within subclade
Confirmed within subclade
Investigational items
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SYMBOLS:
^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for minimum number of reads.
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^^ Indicates an entry which does not meet quality guidelines but may be helpful.
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~ Indicates only an approximate location on the tree.
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The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing done in multiple labs, and/or being found in the area of the chromosome used in recent research studies.
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SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup while regular font items are from Sanger sequencing.
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DID YOU NOTICE that the items in gray sometimes have a long name preceding their listing that is the same as just above? In this situation, evidence seems to indicate they also seem to belong to the same subgroup, but some confirmatory evidence is not yet available.
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EM96/PF1823, A10822/M5550/PF1873, CTS433/M5384/PF1504, CTS860/M5388/PF1511, CTS939/M5389, CTS1008/M5391/PF1516, CTS1105/M5397, CTS2401/M5448/PF1633, CTS2496/M5449/PF1636, CTS2893/M5453, CTS3199/M5455/PF1649, CTS3337/PF1653, CTS3749/M5461/PF1660, CTS3774/M5462/PF1661, CTS4373/M5466/PF1669, CTS4685/M5467/PF1671, CTS4994/M5468, CTS5316/M5470/PF1683, CTS6048/M5475/PF1697, CTS6513/M5483/PF1710, CTS6755/M5485/PF1713, CTS6818/M5487, CTS7138/M5492/PF1723, CTS7824/M5495/PF1732, CTS8053/M5497/PF1738, CTS8189/M5500, CTS8269/M5501/PF1743, CTS8631/L794/M5505/PF1751, CTS8754/M5506/PF1752, CTS9440/M5510/PF1769, CTS9799/M5516/PF1777, CTS10296/M5519/PF1780/V4064, CTS10330/M5520/PF1781/V4089, CTS10344/M5521/PF1783, CTS10894/M5552, CTS11216/M5556/PF1882, CTS11504/M5559/PF1892, CTS11599/PF1893, CTS11622/PF1894, CTS11644/PF1895, CTS11811/M5560/PF1899, CTS11916/M5562/PF1901, FGC2058/S14723/YFS32979/Z15673, FGC2060/Z15671, FGC2079, FGC2647/S7472/Z15672, L339, L504/PF1803, L507/PF1874, L511/PF1888, L537/PF1507, L614/PF1898/YSC0000032, L856/PF1553, M40/PF1445/SRY4064/SRY8299, M5382/PF1502, M5396, M5403/PF1530/V1254, M5406/PF1539, M5410/PF1545/V1692, M5416/PF1552, M5417/PF1554/V2187, M5418/PF1555/V2350, M5422/PF1561/V2471, M5424/PF1563, M5425/PF2323, M5426/PF1564, M5428, M5431/PF1576, M5436/PF1583, M5438, M5439/PF1590, M5440/PF1591, M5452/PF1642, M5525/PF1795, M5527/PF1796, M5529/PF1798, M5533, M5539, M5540/PF1816, M5543/PF1819, M5545/PF1822, M5548/PF1834, M5549/PF1837, M5564/PF1904, M5566/PF1906, M5569/PF1919, M5571/S7474, M5572/PF1923, M9051, M9052, P29/PF1643, P150/PF1704, P152/PF1646, P155/PF1687, P156/PF1725, P168, P169/Page54/PF1878, P170/PF1656, P171/PF1903, P172/PF1510, P173, P174/PF1674, P175, P176/PF1678, PF1567, PF1618, PF1788, PF1868, PF1917
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E1P147/PF1938, CTS955/M5390/PF1514, CTS5913/M5474/PF1694, CTS6355/M5479/PF1703, CTS8432/PF1745, CTS9083/M5509/PF1764, CTS9753/M5515/PF1776, M5413/PF1549/V1844, M5421/PF1559, M5534/PF1809, PF1593
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E1aM132, CTS21, CTS140, CTS245/V1118, CTS340, CTS443, CTS520, CTS548, CTS600, CTS685, CTS895, CTS1059, CTS1076, CTS1134, CTS1172, CTS1364, CTS1374, CTS1404, CTS1671/Z924, CTS1686/Z925, CTS1720, CTS1876, CTS1923, CTS1931, CTS1937/Z926, CTS1987, CTS2002, CTS2031, CTS2131/Z927, CTS2166/Z928, CTS2186, CTS2255, CTS2280, CTS2362, CTS2384/Z929, CTS2743, CTS2751, CTS2808/L633, CTS3045/V2509/Z930, CTS3173, CTS3309/V2623, CTS3342, CTS3459, CTS3507/V2736, CTS3581/V2785, CTS3698, CTS3716, CTS4034, CTS4268, CTS4467/V3085, CTS4477, CTS4591/V3104/Z933, CTS4946, CTS5009/V3271, CTS5033, CTS5194, CTS5283, CTS5313/Z935, CTS5393/Z936, CTS5503/Z937, CTS5589, CTS5701, CTS5734, CTS5775, CTS5793, CTS5905, CTS6063, CTS6213, CTS6230, CTS6245, CTS6288, CTS6370, CTS6410, CTS6476, CTS6486, CTS6516, CTS6608/Z939, CTS6614/Z940, CTS7156, CTS7190, CTS7296, CTS7584, CTS7651, CTS7654, CTS7830, CTS8185, CTS8383, CTS8433, CTS8482, CTS8538, CTS8632, CTS8663, CTS8720, CTS8835, CTS8867, CTS8953, CTS9003/V3320, CTS9074/V3362, CTS9110/V3403, CTS9114/Z942, CTS9155, CTS9198/V3464, CTS9439, CTS9505/Z943, CTS9520, CTS9714, CTS9927, CTS10067, CTS10147, CTS10182/V4005, CTS10475, CTS10497/V4170, CTS10567/V4219, CTS10644/Z957, CTS10752, CTS10964, CTS11187, CTS11248, CTS11276/Z960, CTS11300/Z961, CTS11329, CTS11444, CTS11860, CTS12046/Z962, L481, M10372.1/ZS6696.1, M33, V1277/Z15456, V1341/Z904, V1374/Z15469, V1563/Z15461, V1668/Z15462, V1676/Z15463, V1698/Z15464, V1733/Z15465/Z17396, V1751/Z907, V2008/Z910, V2052/Z15467, V2079/Z911, V2108/Z15468, V2125/Z912, V2310/Z916, Y1193.2, Z913, Z917, Z920, Z921, Z922, Z931, Z938, Z944, Z945, Z946, Z948, Z949, Z950, Z951, Z953, Z963, Z964, Z965, Z966, Z15135, Z15142, Z15454, Z15455, Z15457, Z15458, Z15459, Z15460, Z15466, Z15470, Z15472, Z15473, Z15474, Z15476, Z15478, Z15479, Z15480, Z15481, Z15482, Z15485, Z15486, Z15492, Z15493, Z15494, Z15495, Z15496, Z15499, Z15500/Z17398, Z15501, Z15502, Z15503, Z15504, Z15505, Z15506, Z15507, Z15510, Z15511, Z15512, Z15513, Z15515, Z15516, Z15517/Z17400, Z15518, Z15519, Z15520, Z15521, Z15522, Z15523, Z15524, Z15525, Z15526, Z15527, Z15528, Z15541, Z15542, Z15543, Z15544, Z15545, Z15547, Z15550, Z15551, Z15552, Z15554, Z15556, Z15557, Z17399, Z17403
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E1a1M44, BY172.1/SK563.1/V3797.1/Z17470.1, L632, L634, L1239, L1240, S11513/SK562/V2068/Z17420, SK564/Z17450, SK565/Z17480, SK566/V2316/Z17422, SK567/V1740/Z17417, SK568/Z17498, V1584/Z17412, V1776/Z17418, V1971/Z17419, V2142/Z17421, V2407/Z17423, V2463/Z17424, V2779/Z17439, V2908/Z17441, V2981/Z17442, V3000/Z17433, V3003/Z17444, V3045/Z17445, V3168/Z17446, V3302/Z17466, V3605/Z17468, V3615/Z17469, V3976/Z17472, Z17406, Z17407, Z17408, Z17409, Z17410, Z17411, Z17413, Z17414, Z17416, Z17425, Z17426, Z17427, Z17428, Z17429, Z17430, Z17431, Z17434, Z17435, Z17436, Z17437, Z17438, Z17440, Z17443, Z17447, Z17448, Z17449, Z17451, Z17452, Z17453, Z17454, Z17455, Z17456, Z17457, Z17458, Z17459, Z17460, Z17461, Z17462, Z17463, Z17464, Z17465, Z17471, Z17473, Z17474, Z17475, Z17476, Z17477, Z17478, Z17479, Z17481, Z17482, Z17483, Z17484, Z17485, Z17499, Z17500, Z17501, Z17502, Z17503, Z17504, Z17506, Z17507
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E1a1a
A8384^/Y15944^
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E1a1a1Z17699
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E1a1a1a
Z17697, Z17698, Z17700
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E1a1a1a1
Z42331, Z42332, Z42333, Z42334, Z42335, Z42337, Z42338, Z42588
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E1a1a1a1aZ17696
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E1a1a1a1b~
Z42339, Z42340, Z42341, Z42342, Z42343
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E1a1a1bA7710
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E1a1a1b1~
Z36092, Z36093, Z36094, Z36095, Z36096
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E1a1a1b2~
Z38881, Z38882, Z38883, Z38884, Z38885
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E1a1a2Z17467
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E1a1a2aZ31503
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E1a1a2b
SK569/Z20647, Y20482, Y20483, Y20484, Y20485, Y20486, Y20488, Y20490, Y20646, Y20649, Z40502
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E1a1a2c~
Z20650, Z20651, Z20652, Z20653, Z20654
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E1a1a2d~
Y20219, Y24556, YP5682, YP5685
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E1a1a2d1~
Y24438, Y24439, Y24440, Y24549, Y24550/Z39203, Y24551/Z39204, Y24552, Y24553, Y24554/Z39202, Y24555
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E1a1a2e~
A1688, M3937/S9663, S21504, Y24688, Y24690, Y24693, YP5684, Z20769, Z20771
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E1a1a2e1~
Z20767, Z20770, Z20772, ZS4521.2
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E1a1b~
Z31497, Z31498, Z31500, Z31501
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E1a2CTS10713/Z958, CTS230/V1113, CTS248, CTS678, CTS986, CTS1103, CTS1306, CTS1319, CTS2223, CTS2307, CTS2327, CTS2454, CTS2832, CTS3208, CTS3256, CTS3355/V2643, CTS3515, CTS3538, CTS3914, CTS4174/Z932, CTS4276, CTS4316, CTS4337/V3057, CTS4692, CTS5155/Z934, CTS5422, CTS5458, CTS6329, CTS6384, CTS6891, CTS7453/Z941, CTS7499, CTS7978, CTS8133, CTS8817, CTS9096/V3384, CTS9548/V3633, CTS10225, CTS10801, CTS11085/Z959, CTS11320, CTS11390, CTS12022, V1266/Z15127, V1507/Z905, V2232/Z915, V1312/Z15128, Z908, Z914, Z955, Z956, Z967, Z15129, Z15130, Z15131, V2002/Z15132, Z15133, Z15134, Z15136, Z15141, Z15145, Z15148, Z15151, Z15152, Z15153, Z15155, Z15162, Z15163, Z15165, Z15169
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E1a2~
CTS6817, CTS7198
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E1a2a
CTS246/V1119, CTS2632, CTS4519, CTS9089/V3379, Y967, Z906, Z918, Z15171
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E1a2a1
CTS10935, Z15271
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E1a2a1a
CTS3380, CTS2648, CTS4187, V1734/Z885
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E1a2a1a1P110, CTS1753, CTS1927/Z888, CTS2273, CTS2518/Z889, CTS3519, CTS3881/V2847, CTS5110^/Z890^, CTS5365^, CTS7182^, CTS8725^, CTS11593, CTS11290^/Z894^, CTS11578/Z895, V2058/Z886, Z887, Z892^, Z15395, Z15396^
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E1a2a1a1aCTS1792, CTS6482, CTS7040, CTS7797, CTS10874, V4008/Z15403, Z15397, Z15398, Z15399, Z15400, Z15401, Z15402, Z15404, Z15405
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E1a2a1a1a1~
Z36289, Z36290, Z45457, Z45458, Z45459
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E1a2a1a1a2~
CTS1341, CTS10757, Z36286, Z36287
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E1a2a1a1b~
CTS5362, CTS5732, CTS7020, CTS8238, CTS9043
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E1a2a1a2
Z15407, Z15408, Z15409, Z15410
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E1a2a1a2~
Z15406
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E1a2a1a2a~
Z36296, Z36297, Z36298, Z36299
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E1a2a1a2b~
Z36300, Z36301, Z36302, Z36303, Z36304
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E1a2a1bL133.1/Page74.1, CTS1285, CTS5145/Z880, CTS5607, CTS6573, CTS8762/Z881, CTS11769, Z878, Z879, Z15274, Z15276, Z15277, Z15278, Z15312
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E1a2a1b1CTS736, A173.1/Z15291.1, CTS602, CTS3351, CTS3615, CTS3840, CTS7084, CTS7140, CTS7912, CTS9489/M1146, CTS9792, CTS10712, CTS11737, Z12510.2/Z15303.2, Z15285, Z15286^, Z15287, Z15288, Z15289, Z15290, Z15292, Z15293, Z15294, Z15299, Z15300, Z15301, Z15302, Z15304, Z15305^, Z15306/ZS6189, Z15308, Z15309, Z15310, Z15311, Z15312, Z15313, Z15320, Z15322, Z15323, Z15324
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E1a2a1b1~
CTS7084, CTS9489/M1146, Z15320, Z15323
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E1a2a1b1aCTS3764.1^, FGC24648/Z15361, Z5992, Z15339, Z15340, Z15341, Z15342, Z15343, Z15344, Z15345, Z15346, Z15348, Z15349, Z15350, Z15351, Z15352, Z15353, Z15354, Z15355, Z15356, Z15358, Z15359, Z15360^, Z15362, Z15363^, Z15364, Z15365, Z15366, Z15367, Z15368, Z15369, Z15371^, Z15372, Z15373, Z15374, Z15375, Z15376, Z15377, Z15378, Z15379, Z15380, Z15381^, Z15382, Z15383, Z15384, Z15391, Z15392, Z15393^, Z15394
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E1a2a1b1a1~
Z36305, Z36306, Z36308, Z36309
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E1a2a1b1a2~
Z42529, Z42531, Z43245
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E1a2a1b1b~
CTS2361, CTS3491, CTS4038, CTS6669, CTS7911
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E1a2a1b2~
CTS749, CTS2333, CTS3095, CTS4106, CTS4315
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E1a2a1b3~
L262
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E1a2a1c~
Z36237, Z36238, Z36239, Z36240, Z36288
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E1a2a2Z5991, FGC23407/Z15201, Z15172, Z15183, Z15184, Z15185, Z15186, Z15187, Z15188^, Z15189, Z15190, Z15191, Z15192, Z15193, Z15194, Z15195, Z15196, Z15197, Z15198, Z15199, Z15200, Z15204, Z15205, Z15207, Z15208, Z15212, Z15217, Z15218, Z15219, Z15220, Z15221, Z15222, Z15223, Z15224, Z15225, Z15226, Z15227, Z15228, Z15229, Z15230, Z15231, Z15232, Z15233, Z15234, Z15235, Z15236, Z15237^, Z15238, Z15239, Z15240^, Z15241, Z15242, Z15244, Z15245, Z15246, Z15247, Z15249, Z15250, Z15252, Z15253, Z15254^, Z15255, Z15256, Z15257, Z15259, Z15260, Z15261, Z15262, Z15263, Z15265, Z15266, Z15268, Z15269, Z15270, Z20658, Z20722^, Z20734, Z20747^
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E1a2a2a~
Z36227, Z36229, Z36231, Z42534, Z43246
76
E1a2a2b~
Z36232, Z36233, Z36234, Z36235, Z36236
77
E1a2b
Z5985.1, Z15088, Z15089, Z15090, Z15092, Z15093, Z15096, Z15097, Z15098, Z15099, Z15102, Z15103, Z15104
78
E1a2b~
Z15094, Z15095
79
E1a2b1
L94, Z15068, Z15070, Z15071, Z15072, Z15073, Z15075, Z15076, Z15077, Z15078, Z15079, Z15080, Z15082, Z15083
80
E1a2b1a
Z15064, Z15065, Z15067
81
E1a2b1a1Z5988
82
E1a2b1a1aZ14979
83
E1a2b1a1a1Z14889, FGC13367/Z14956, Z14898, Z14899, Z14900, Z14901, Z14902, Z14903, Z14904, Z14905, Z14906, Z14907, Z14908, Z14909, Z14910, Z14911, Z14912, Z14913, Z14914, Z14915, Z14916, Z14917, Z14918, Z14919, Z14923, Z14925, Z14926, Z14928, Z14930, Z14931, Z14932, Z14933, Z14934, Z14935, Z14937, Z14938, Z14939, Z14940, Z14941, Z14942, Z14943, Z14944, Z14945, Z14946, Z14947, Z14948, Z14949, Z14950, Z14951, Z14952, Z14953, Z14954, Z14955, Z14957, Z14958, Z14959, Z14967, Z14968, Z14969, Z14970, Z14971, Z14972, Z14973
84
E1a2b1a1a1~
Z14936
85
E1a2b1a1a1a~
Z36345, Z36346, Z36347, Z36348, Z36349
86
E1a2b1a1a1b~
Z36350, Z42536, Z42537
87
E1a2b1a1a1c~
Z36352, Z36353, Z36354, Z36355, Z36356
88
E1a2b1a1a2~
CTS11417.2/M2288.2, Z36341, Z36342, Z36343, Z36344
89
E1a2b1a2Z5987, Z14992, Z14994, Z14995, Z14996, Z14997, Z14998, Z14999, Z15000, Z15002, Z15007, Z15008, Z15009, Z15010, Z15011, Z15012, Z15014, Z15015, Z15018, Z15019, Z15020, Z15021, Z15022, Z15023, Z15024, Z15025, Z15027, Z15028, Z15029^, Z15030, Z15031, Z15032, Z15033, Z15034, Z15035, Z15036, Z15038, Z15040, Z15041, Z15042, Z15043, Z15044, Z15045, Z15049, Z15050, Z15051, Z15052, Z15053, Z15054, Z15055.1, Z15056, Z15059, Z15061, Z15170
90
E1a2b1a2a~
Z36319, Z36320, Z36321, Z36322
91
E1a2b1a2b~
Z36324, Z36325, Z36326, Z36327, Z36328
92
E1a2b1b~
Z36314, Z36315, Z36317, Z36318
93
E1a2b1c~
PF3664.2/V2273.2/Z2638.2
94
E1a2c~
Z36222, Z36223, Z36224, Z36225, Z36226
95
E1a3~Z322
96
E1b
P177/PF1939
97
E1b1
P2/PF1940/PN2, P179, P180/PF1941, P181
98
E1b1~CTS119/M5377/PF1448/V1048, CTS331/M5383/PF1503, CTS1446/P178/PF1526, CTS1867/M5443/PF1624, CTS1978/M5445/PF1626, CTS2150/M5446/PF1628, CTS2631/M5450, CTS2670/M5451/PF1639, CTS3250/M5456/PF1650, CTS3262/M5457/PF1651/V2598, CTS3362/M5460/PF1654, CTS4852, CTS5373/M5471/PF1684, CTS5404/M5472/PF1685, CTS6232/M5477, CTS6333^/M5478^/PF1702^, CTS6707/M5484/PF1712, CTS6842/M5488/PF1719, CTS7058/M5490/PF1722, CTS7133/M5491, CTS7364/M5493/PF1726, CTS7913/M5496/PF1734, CTS8159/M5499/PF1742, CTS8369/M5502/PF1744, CTS8498/M5503/PF1747, CTS8795/M5507/PF1753, CTS8900/PF1756, CTS8947/M5508/PF1760/V3292, CTS9573/M5512, CTS10046/M5517, CTS10385/M5522/PF1885, CTS12006/M5563/PF1902, F1365/M5420/PF1558, FGC2070, L487/PF1589, L489/M5444/PF1625, L492/M5463/PF1667, L499/M5504/PF1750, L510/M5555/PF1880, L512/M5561/PF1900, L535/M5401, L613/M5437/PF1584, M5404/PF1535, M5405/PF1536, M5407/PF1541, M5409/PF1544, M5411/PF1547, M5412/PF1548/V1798, M5415/PF1551, M5423^/PF1562^/V2484^, M5429/PF1565, M5432/PF1577, M5433/PF1578, M5441, M5481/PF1709, M5524/PF1792, M5530/PF1799, M5536/PF1811, M5544/PF1821, M5546/PF1827, M5547/PF1828, M5570/PF1920, PF1610, PF1838
99
E1b1a
TSC0077541/V38
100
E1b1a1
M291, P1/PN1, P189.1, P293.1, Page106, CTS5964/TSC0424987, Z1101, Z1116, Z1123, Z1125, Z1127, Z1130, Z1133, Z1135
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