ACDEFGHIJKLMNOPQRSTUVWXYZAAABACADAEAFAGAHAIAJAKALAMANAOAPAQARASATAUAVAWAXAYAZBABBBCBDBEBFBGBH
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Catalogue of the Portuguese Bryophytes (2003)
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Adapted from: Sérgio, C. & Carvalho, S. (2003). Annotated Catalogue of Portuguese Bryophytes. Portugaliae Acta Biol. 21: 5-A3
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NUMBER OF OBSERVED TAXA (Portugal): 527
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BRYOPHYTA (Species Names)Regiões ("Províncias")Google/SearchGoogle/Fotos
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Acaulon fontiquerianum Casas & SérgioMedmap: Cogoni & Scrugli, 2000.
IPmap: Casas et al., 1989.
E: Casas et al., 1989; Sérgio et al., 1993a.		Acaulon fontiquerianum Acaulon fontiquerianum
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Acaulon muticum (Hedw.) Müll. Hal. var. muticumAg: Solms-Laubach, 1868; Henriques, 1889; Machado, 1928; Casares, 1932; Sérgio
et al., 1984; Sérgio et al., 1987a.
BL: Sérgio et al., 1988.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Machado, 1928;
Casares, 1932.
Pt: Sérgio et al., 1987a.		Acaulon muticum Acaulon muticum
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Acaulon muticum (Hedw.) Müll. Hal. var. mediterraneum (Limpr.) SérgioPtmap: Sérgio, 1972c.
Ag: Sérgio, 1972c; Casas et al., 1990.
BAl: Sérgio, 1972c.
BB: Sérgio et al., 1994d, Sérgio et al., 2002c.
E: Sérgio, 1972c.
R: Sérgio et al., 1994d.		Acaulon muticum Acaulon muticum
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Acaulon triquetrum (Spruce) Müll. Hal.IPmap: Casas et al., 1989.
Ag: Sérgio et al., 1984; Sérgio et al., 1987a; Casas et al., 1989.		Acaulon triquetrum Acaulon triquetrum
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Aloina aloides (Schultz) Kindb.AAl: Sérgio et al., 1997b. E: Mitten, 1853; Henriques, 1889; Luisier, 1916b; Pereira-Coutinho, 1917b;
Machado, 1928; Delgadillo, 1975; Sérgio & Sim-Sim, 1985a; Galego et al.,1999;
Galego et al., 2000.
Mi: Machado, 1920a; Machado, 1928; Sérgio & Schumacker, 1992.
R: Sérgio et al., 1988.
TM: Machado, 1915b.
Pt: Sérgio et al., 1987a.
Ag: Crundwell, 1956; Sérgio et al., 1984; Sérgio et al., 1987a.
BA/BB: Greven & Melick, 1994.
BL: Henriques, 1889; Machado, 1928; Sérgio et al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1928.		Aloina aloides Aloina aloides
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Aloina ambigua (Bruch & Schimp.) Limpr.AAl: Luisier, 1910c; Luisier, 1916b; Machado, 1928; Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Pereira-Coutinho, 1917b; Delgadillo,
1975; Sérgio et al., 1984; Sérgio et al., 1987a; Galego et al.,1999; Galego et al.,
2000.
BL: Brotero, 1804; Henriques, 1889; Anónimo, 1893; Machado, 1928; Sérgio et al.,
1988.
DL: Henriques, 1889; Machado, 1928.
E: Brotero, 1804; Mitten, 1853; Henriques, 1889; Luisier, 1907a; Luisier, 1910c;
Luisier, 1916b; Pereira-Coutinho, 1917b; Machado, 1928; Ade & Koppe, 1942;
Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985a.
Mi: Machado, 1913; Machado, 1916; Machado, 1920a; Machado, 1928; Sá-
Nogueira, 1950d; Sérgio & Schumacker, 1992.
R: Pereira-Coutinho, 1917b; Machado, 1928; Sérgio et al., 1988.
Pt: Sérgio et al., 1987a.		Aloina ambigua Aloina ambigua
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Aloina rigida (Hedw.) Limpr.Ag: Galego et al., 2000.Aloina rigida Aloina rigida
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Amblystegium humile (P. Beauv.) Crundw.Ag: Sérgio et al., 1984.Amblystegium humile Amblystegium humile
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Amblystegium riparium (Hedw.) Schimp.AAl: Sérgio et al., 1997b.
Ag: Dixon, 1912; Machado, 1933; Allorge, 1935; Allorge, 1974; Sérgio et al., 1984.
BA: Pierrot, 1972.
BAl: Sérgio, 1970-1971; Allorge, 1974.
BB: Luisier, 1916b; Pereira-Coutinho, 1917b; Machado, 1933; Allorge, 1974; Sérgio
et al., 2002c.
BL: Henriques, 1889; Pereira-Coutinho, 1917b; Machado, 1933; Allorge, 1974;
Sérgio et al., 1988; Leitão et al., 1996; Werner, 1997.
DL: Anónimo, 1883a; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Pereira-Coutinho, 1917b; Machado, 1933; Allorge, 1974; Werner, 1997.
E: Luisier, 1916b; Pereira-Coutinho, 1917b; Luisier, 1924; Machado, 1933; Allorge,
1974; Sérgio & Sim-Sim, 1985a.
Mi: Brotero, 1804; Henriques, 1889; Machado, 1913; Luisier, 1924; Machado, 1933;
Sá-Nogueira, 1950d; Allorge, 1974; Sérgio & Schumacker, 1992.
R: Sérgio et al., 1988; Ferreira, 1992.
TM: Allorge, 1974.
Pt: Luisier, 1924.		Amblystegium riparium Amblystegium riparium
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Amblystegium serpens (Hedw.) Schimp.Ag: Dixon, 1912; Machado, 1933; Sérgio, 1970-1971.
[BA/BB: Greven & Melick, 1994].
BL: Brotero, 1804; Machado, 1933; Sérgio, 2001b.
The more recent report is to Serra da Estrela (BA/BB), however this species, not being
confirmed by any herbarium specimens, is to be considered as uncertain.		Amblystegium serpens Amblystegium serpens
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Amblystegium varium (Hedw.) Lindb.BA: Sérgio et al., 2001a.Amblystegium varium Amblystegium varium
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Amphidium lapponicum (Hedw.) Schimp.[Mi: Luisier, 1948].
This sub-arctic species was only reported from one locality from Minho by Luisier.
However, the report was based on the specimen recently named as Bryoerythrophyllum
campylocarpum by Sérgio (1992).		Amphidium lapponicum Amphidium lapponicum
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Amphidium mougeotii (Bruch & Schimp.) Schimp.IPmap: Casas et al., 1989.
BA: Casas et al., 1989; Sérgio, 1994a; Sérgio et al., 2001d.
BA/BB: Greven & Melick, 1994.
BL: Casas et al., 1989.
Mi: Machado, 1917a; Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1928;
Casares, 1932; Sá-Nogueira, 1950c; Sá-Nogueira, 1950d; Casas et al., 1989; Sérgio
& Schumacker, 1992.
TM: Casas et al., 1989; Sérgio & Schumacker, 1992.		Amphidium mougeotii Amphidium mougeotii
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Anacolia webbii (Mont.) Schimp.Medmap: Cogoni et al., 1999.
IPmap: Casas et al., 1985.
BA: Casas et al., 1985.
TM: Machado, 1930; Allorge, 1931a; Machado, 1933; Allorge & Allorge, 1949;
Flowers, 1952; Casas et al., 1985.		Anacolia webbii Anacolia webbii
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Andreaea crassinervia Bruch[BA: Henriques, 1883; Henriques, 1889; Machado, 1917b; Pereira-Coutinho, 1917;
Machado, 1933; Sérgio, 1994a; Sérgio & Séneca, 1994.
BA/BB: Levier, 1880; Greven & Melick, 1994.
BB: Machado, 1933; Sérgio & Séneca, 1994].
The revision of the specimens named as Andreaea crassinervia revealed that most of the
material corresponds to A. rothii or A. heinemannii subsp. heinemannii (Sérgio, in press).
So, in Portugal and perhaps even in the Iberian area, the species A. crassinervia is
certainly not present (Murray comm.).		Andreaea crassinervia Andreaea crassinervia
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Andreaea frigida HuebenerIPmap: Sérgio et al., 1994c; Casas et al., 1996 ; Sérgio & Draper, 2002b.
BA: Casas et al., 1996; Sérgio et al., 2001e; Sérgio et al., 2002a.
BA/BB: Casas et al., 1996.
Pt: Murray, 1988. Recorded only in Serra da Estrela it is considered extinct (Sérgio et al., 1994). However, was recently discovered in different localities in the same mountain range (Sérgio et al., 2002a).		Andreaea frigida Andreaea frigida
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Andreaea heinemannii Hampe & Müll. Hal. subsp. heinemanniiWmap: Schultze-Motel, 1970b.
See note of A. crassinervia.		Andreaea heinemannii Andreaea heinemannii
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*Andreaea heinemannii Hampe & Müll. Hal. subsp. crassifolia (Luisier) SérgioIPmap: Casas et al., 1996.
BA: Pereira-Coutinho, 1917b; Sérgio & Séneca, 1994; Casas et al., 1996.
BA/BB: Schultze-Motel, 1970a.
BB: Luisier, 1916a; Luisier, 1916b; Casares, 1932; Machado, 1933; Schultze-Motel,
1970b; Sérgio & Séneca, 1994; Casas et al., 1996.
DL: Sérgio, 2003 (Not Published).
Mi: Sérgio & Séneca, 1994; Casas et al., 1996.
TM: Casas et al., 1996.
Pt: Luisier, 1934.
Records from Portugal originally thought to be of Andreaea crassifolia Luisier have been
redetermined as A. heinemannii subsp. crassifolia (Luisier) Sérgio (Sérgio, in press). With
a South Atlantic/Macaronesian distribution, all the reports of A. heinemannii (Casas et al.,
1996) correspond to this subspecies.		*Andreaea heinemannii *Andreaea heinemannii
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Andreaea megistospora B. M. MurrayIPmap: Casas et al., 1996.
DL: Sérgio & Séneca, 1994; Casas et al., 1996.
Mi: Sérgio & Schumacker, 1992; Sérgio & Séneca, 1994; Casas et al., 1996.
TM: Sérgio & Schumacker, 1992; Casas et al., 1996.		Andreaea megistospora Andreaea megistospora
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Andreaea rothii F. Weber & D. Mohr s. l.Eurmap: Schultze-Motel, 1970a.
IPmap: Casas et al., 1996.
AAl: Casas et al., 1996.
BA: Machado, 1933; Sérgio, 1994a; Casas et al., 1996.
BA/BB: Greven & Melick, 1994; Casas et al., 1996.
BB: Casas et al., 1996.
BL: Casas et al., 1996.
DL: Casas et al., 1996; Werner, 1997.
Mi: Reinoso et al., 1990; Sérgio & Schumacker, 1992; Casas et al., 1996.
TM: Casas et al., 1996.
The identification of the subspecies of the lastest reports still needs to be recognized.		Andreaea rothii Andreaea rothii
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Andreaea rothii F. Weber & D. Mohr subsp. rothiiAAl: Sérgio et al., 1997b.
BA: Luisier, 1916b; Pereira-Coutinho, 1917b.
BA/BB: Machado, 1917b; Pereira-Coutinho, 1917b.
BB: Luisier, 1916b; Machado, 1933.
Mi: Mitten, 1853; Henriques, 1889; Luisier, 1907; Machado, 1915a; Luisier, 1916b;
Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1920a; Machado, 1933; Sá-
Nogueira, 1950c; Sá-Nogueira, 1950d; Sérgio & Schumacker, 1992.
TM: Ervideira, 1926.
Pt: Schultze-Motel, 1970a; Machado, 1933.		Andreaea rothii Andreaea rothii
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Andreaea rothii F. Weber & D. Mohr subsp. falcata (Schimp.) Lindb.BA: Henriques, 1883; Henriques, 1889; Schultze-Motel, 1970a.
BA/BB: Casares, 1932.
BB: Casares, 1932.
Mi: Machado, 1920a; Casares, 1932; Sá-Nogueira, 1950c; Sá-Nogueira, 1950d;
Sérgio & Schumacker, 1992.
TM: Ervideira, 1922; Sérgio & Schumacker, 1992.
Pt: Machado, 1933; Casares, 1932.		Andreaea rothii Andreaea rothii
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Andreaea rupestris Hedw.IPmap: Casas et al., 1996.
BA: Sérgio, 1994a; Sérgio & Séneca, 1994; Casas et al., 1996.
BA/BB: Levier, 1880; Henriques, 1883; Casares, 1932; Casas et al., 1996.
BB: Machado, 1933.
Mi: Anónimo, 1887; Anónimo, 1890; Sampaio, 1901-1902; Machado, 1913;
Machado, 1913; Machado, 1917b; Machado, 1933; Casares, 1932; Sá-Nogueira,
1950d; Sérgio & Schumaker, 1992; Casas et al., 1996.
TM: Sérgio & Schumaker, 1992; Casas et al., 1996.		Andreaea rupestris Andreaea rupestris
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Anomobryum julaceum (P. Gaertn., B. Mey. & Scherb.) Schimp.AAl: Allorge, 1974.
Ag: Solms-Laubach, 1868 original ; Welwitsch, 1872; Henriques, 1889; Luisier,
1910c; Dixon, 1912; Pereira-Coutinho, 1917b; Luisier, 1924; Machado, 1930;
Loeske, 1932; Allorge, 1935; Sérgio, 1970-1971; Pierrot, 1972; Allorge, 1974;
Guerra et al., 1981; Jiménez et al., 2002.
BA: Henriques, 1883, Henriques, 1889; Pais, 1964; Allorge, 1974; Sérgio, 1994a.
BA/BB: Pereira-Coutinho, 1917b; Machado, 1930; Greven & Melick, 1994.
BAl: Sérgio et al.,1999a; Sérgio & Garcia, 2001b.
BB: Luisier, 1910c; Luisier, 1916b; Pereira-Coutinho, 1917b.
BL: Warnstorf, 1899; Allorge, 1935; Allorge, 1974.
DL: Henriques, 1889; Sampaio, 1901-1902; Pereira-Coutinho, 1917b.
Mi: Machado, 1913; Machado, 1920a; Barros, 1942b; Barros, 1944b; Sá-Nogueira,
1950d; Allorge, 1974; Sérgio & Schumacker, 1992.
TM: Machado, 1915b; Ervideira, 1922.
The original description as Anomobryum juliforme in Solms-Laubach (1868).		Anomobryum julaceum Anomobryum julaceum
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*Anomobryum lusitanicum (I. Hagen in Luisier) Thér.IPmap: Sérgio et al., 1994c; Casas et al., 1996; Sérgio & Draper, 2002b.
AAl: Allorge, 1935; Pierrot, 1972; Casas et al., 1996; Sérgio et al., 1996; Sérgio et
al., 1997b.
BA: Allorge, 1931b; Machado, 1933; Casas et al., 1996; Sérgio et al., 1996.
BB: Luisier, 1915a (Orig. descrip.), Luisier, 1915c; Luisier, 1915d; Luisier, 1916b;
Luisier, 1924; Machado, 1930; Allorge, 1931b; Casas et al., 1996; Sérgio et al.,
1996.
Mi: Luisier, 1915c.
Pt: Thériot, 1933; Luisier, 1934.
For a review of this Iberian species see Sérgio et al. (1994c).
AAl: Allorge, 1935; Pierrot, 1972; Casas et al., 1996; Sérgio et al., 1996; Sérgio et
al., 1997b.
BA: Allorge, 1931b; Machado, 1933; Casas et al., 1996; Sérgio et al., 1996.		*Anomobryum lusitanicum *Anomobryum lusitanicum
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Anomodon viticulosus (Hedw.) Hook. & TaylorEurmap: Granzow De La Cerda, 1988.
BL: Allorge, 1928a; Machado, 1933; Allorge, 1974; Granzow De La Cerda, 1988;
Sérgio et al., 2002b.
Species only collected in Buçaco by P. & V. Allorge, 1928 and 1929. It was considered to
have disappeared from the area (Allorge, 1974), however there is a recent gathering in
Serra de Sicó (Sérgio et al., 2002b).		Anomodon viticulosus Anomodon viticulosus
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Antitrichia californica Sull.Wmap: Allorge, 1931c; Sérgio, 1990.
IPmap: Casas et al., 1985.
Ptmap: Sim-Sim & Sérgio, 1998.
AAl: Sá-Nogueira, 1958; Sérgio, 1969; Pierrot, 1972; Casas et al., 1985; Sérgio et
al., 1997b.
AAl/BAl: Allorge, 1938.
BA: Allorge, 1931b; Machado, 1933; Sérgio, 1969; Pierrot, 1972; Casas et al., 1985;
Sérgio, 1994a.
BB: Allorge, 1938; Casas et al., 1985; Catarino et al., 1987.
R: Catarino et al., 1987; Sérgio, C. & Sim-Sim, M., 1994; Sérgio et al., 1998d.
TM: Allorge, 1931b; Machado, 1933; Machado, 1933; Allorge, 1938; Allorge &
Allorge, 1949; Casas et al., 1985.		Antitrichia californica Antitrichia californica
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Antitrichia curtipendula (Hedw.) Brid.IPmap: Casas et al., 1989.
Ptmap: Sim-Sim & Sérgio, 1998.
AAl: Casas et al., 1989.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Dixon, 1912;
Luisier, 1924; Machado, 1933; Sérgio, 1970-1971; Allorge, 1974; Casas et al., 1989.
BA: Henriques, 1883, Henriques, 1889; Pereira-Coutinho, 1917b; Allorge, 1974;
Casas et al., 1989; Jansen, 1994; Sérgio, 1994a; Sérgio et al., 2001d.
BA/BB: Machado, 1917b; Pereira-Coutinho, 1917b; Greven & Melick, 1994.
BB: Pereira-Coutinho, 1917b; Machado, 1933; Allorge, 1974; Casas et al., 1989.
BL: Henriques, 1889; Machado, 1933; Allorge, 1974; Casas et al., 1989.
DL: Casas et al., 1989.
E: Brotero, 1804; Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b;
Machado, 1933; Allorge, 1974; Casas et al., 1989.
Mi: Brotero, 1804; Henriques, 1889; Sampaio, 1901-1902; Machado, 1913;
Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1920a; Luisier, 1924;
Machado, 1933; Barros, 1944b; Sá-Nogueira, 1950d; Allorge, 1974; Casas et al.,
1989; Sérgio & Schumacker, 1992; Albertos, 2001; Calleja et al., 2001.
TM: Ervideira, 1922; Ervideira, 1926; Machado, 1933; Allorge & Allorge, 1949;
Allorge, 1974; Casas et al., 1989; Sérgio & Schumacker, 1992.		Antitrichia curtipendula Antitrichia curtipendula
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Archidium alternifolium (Hedw.) Mitt.Wmap: Sérgio, 1990.
IPmap: Casas et al., 1992.
AAl: Sérgio et al., 1997b; Sérgio et al., 1998c.
Ag: Solms-Laubach, 1868; Henriques, 1889; Machado, 1928; Casares, 1932; Pierrot,
1972; Sérgio et al., 1984; Sérgio, 1991; Casas et al., 1992.
BA: Sérgio, 1991; Casas et al., 1992; Jansen & Sequeira, 1999.
BAl: Sérgio, 1991; Casas et al., 1992; Sérgio et al., 1998c; Sérgio et al.,1999a.
BB: Sérgio et al., 1998c; Sérgio et al., 2002c.
BL: Sérgio et al., 1988; Casas et al., 1992.
E: Sérgio, 1972a; Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985a; Sérgio et al.,
1988; Arts, 1990; Sérgio, 1991; Casas et al., 1992.
R: Sérgio et al., 1988; Sérgio, 1991; Casas et al., 1992; Sérgio et al., 1998c.		Archidium alternifolium Archidium alternifolium
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Aschisma carniolicum (F. Weber & D. Mohr) Lindb.IPmap: Casas et al., 1985; Gil & Castro, 1987.
Ptmap: Sérgio, 1972c.
AAl: Sérgio et al., 1997b.
BAl: Casas et al., 1985.
BL: Sérgio et al., 1988.
E: Sérgio, 1972c; Sérgio & Correia, 1981; Casas et al., 1985; Sérgio & Sim-Sim,
1985a; Gil & Castro, 1987; Sérgio et al., 1988.		Aschisma carniolicum Aschisma carniolicum
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Astomum crispum (Hedw.) HampeAg: Solms-Laubach, 1868; Henriques, 1889; Machado, 1928; Casares, 1932; Sérgio
et al., 1984; Sérgio et al., 1987a.
BL: Sérgio et al., 1988; Guerra et al., 2002.
DL: Sampaio, 1901-1902; Machado, 1928.
E: Pierrot, 1972; Sérgio & Correia, 1981.
Mi: Luisier, 1907a; Machado, 1913; Machado, 1928; Casares, 1932.
R: Sérgio et al., 1988.
Pt: Sérgio et al., 1987a.		Astomum crispum Astomum crispum
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Astomum levieri (Limpr.) Kindb.Ag: Sérgio et al., 1984; Sérgio et al., 1987a; Guerra et al., 2002.Astomum levieri Astomum levieri
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Atrichum angustatum (Brid.) Bruch & Schimp. var. angustatumAAl: Sérgio et al., 1997b.
Ag: Allorge, 1974.
BL: Allorge, 1931a; Anónimo, 1967; Allorge, 1974.
DL: Henriques, 1889; Sampaio, 1901-1902; Pereira-Coutinho, 1917b; Machado,
1933; Allorge, 1974.
Mi: Machado, 1914a; Luisier, 1916b; Machado, 1920a; Machado, 1933; Allorge,
1974; Sérgio & Schumacker, 1992.
Pt: Luisier, 1924.		Atrichum angustatum Atrichum angustatum
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Atrichum angustatum (Brid.) Bruch & Schimp. var. rhystophyllum (Müll. Hal.) P. W. Richards & E. C. WallaceAg: Dixon, 1912; Machado, 1933; Allorge, 1974.
BL: Dixon, 1912; Machado, 1933; Allorge, 1974.
When no var. is specified in the different reports, we assume that it corresponds to var.
type. However, we think that var. rhystophyllum can be the most frequent taxon in
Portugal.		Atrichum angustatum Atrichum angustatum
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Atrichum undulatum (Hedw.) P. Beauv.Ag: Estácio da Veiga, 1869; Henriques, 1889; Allorge, 1935.
BA: Tavares, 1956; Sérgio, 1969; Allorge, 1974; Sérgio, 1994a; Sérgio et al., 2001d.
BA/BB: Greven & Melick, 1994.
BB: Luisier, 1916b; Pereira-Coutinho, 1917b; Sérgio et al., 2002c.
BL: Henriques, 1889; Warnstorf, 1899; Anónimo, 1969; Allorge, 1974.
DL: Anónimo, 1883a; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Luisier, 1916b; Pereira-Coutinho, 1917b.
E: Henriques, 1889.
Mi: Sampaio, 1901-1902; Machado, 1913; Machado, 1917a; Machado, 1920a;
Machado, 1933; Barros, 1944b; Sá-Nogueira, 1950d; Allorge, 1974; Sérgio &
Schumacker, 1992.
TM: Ervideira, 1922; Sérgio & Schumacker, 1992.
Pt: Machado, 1933.		Atrichum undulatum Atrichum undulatum
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Aulacomnium androgynum (Hedw.) Schwägr.IPmap: Casas et al., 1989.
AAl: Pierrot, 1972; Allorge, 1974; Casas et al., 1989; Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Pereira-Coutinho, 1917b; Machado, 1930; Allorge, 1974;
Casas et al., 1989.
BA: Braun-Blanquet et al., 1956; Casas et al., 1989; Sérgio, 1994a; Jansen &
Sequeira, 1999.
BA/BB: Greven & Melick, 1994.
BB: Allorge, 1974; Jansen, 1994.
BL: Allorge, 1935; Sá-Nogueira, 1950a; Sá-Nogueira, 1950e; Allorge, 1974; Casas
et al., 1989.
E: Barros, 1944a; Sá-Nogueira, 1950e; Allorge, 1974; Casas et al., 1989.
Mi: Henriques, 1889; Machado, 1917b; Machado, 1930; Sá-Nogueira, 1950c; Sá-
Nogueira, 1950d; Allorge, 1974; Casas et al., 1989.
R: Pereira-Coutinho, 1917b; Machado, 1930; Allorge, 1974; Casas et al., 1989.
TM: Ervideira, 1922; Machado, 1930; Allorge & Allorge, 1949; Allorge, 1974;
Casas et al., 1989; Sérgio & Schumacker, 1992.		Aulacomnium androgynum Aulacomnium androgynum
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Aulacomnium palustre (Hedw.) Schwägr.IPmap: Casas et al., 1989.
BA: Henriques, 1883, Henriques, 1889; Machado, 1917b; Pereira-Coutinho, 1917b;
Braun-Blanquet et al., 1952; Tavares, 1956; Casas et al., 1989; Jansen, 1994; Sérgio,
1994a; Jansen & Sequeira, 1999.
BA/BB: Levier, 1880; Sampaio, 1901-1902; Pereira-Coutinho, 1917b; Machado,
1930; Greven & Melick, 1994.
BB: Luisier, 1916b; Casas et al., 1989; Jansen, 1994.
BL: Casas et al., 1989.
DL: Henriques, 1889.
Mi: Pereira-Coutinho, 1917b; Machado, 1920a; Machado, 1930; Sá-Nogueira,
1950c; Sá-Nogueira, 1950d; Teles, 1970; Casas et al., 1989; Sérgio & Schumacker,
1992.
TM: Ervideira, 1922; Machado, 1930; Allorge & Allorge, 1949; Casas et al., 1989;
Sérgio & Schumacker, 1992.
Pt: Luisier, 1924.		Aulacomnium palustre Aulacomnium palustre
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Barbula convoluta Hedw. var. convolutaAAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Pereira-Coutinho, 1917b; Machado,
1928; Sérgio, 1970-1971; Allorge, 1974; Sérgio et al., 1984.
BA: Jansen & Sequeira, 1999.
BL: Sá-Nogueira, 1950a; Sérgio, 1969; Anónimo, 1969; Allorge, 1974; Sérgio et al.,
1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1928; Casares, 1932; Allorge,
1974.
E: Mitten, 1853; Henriques, 1889; Luisier, 1916b; Pereira-Coutinho, 1917b;
Machado, 1928; Casares, 1932; Ade & Koppe, 1942; Allorge, 1974; Sérgio &
Correia, 1981; Sérgio & Sim-Sim, 1985a; Sérgio et al., 1988.
Mi: Machado, 1913; Machado, 1920a; Machado, 1928; Casares, 1932; Barros,
1942b; Sá-Nogueira, 1950d; Allorge, 1974; Sérgio & Schumacker, 1992.
R: Pereira-Coutinho, 1917b; Machado, 1928; Casares, 1932; Sérgio, 1972a; Allorge,
1974; Sérgio et al., 1988.		Barbula convoluta Barbula convoluta
44
Barbula convoluta Hedw. var. commutata (Jur.) Husn.Mi: Machado, 1916; Machado, 1918; Machado, 1920a; Machado, 1928.
The distribution of var. commutata is probably not fully known.		Barbula convoluta Barbula convoluta
45
Barbula ehrenbergii (Lorentz) M. Fleisch.IPmap: Casas et al., 1989; Sérgio et al., 1994c.
E: Pereira-Coutinho, 1917b; Machado, 1928; Allorge, 1931a; Casas et al., 1989.		Barbula ehrenbergii Barbula ehrenbergii
46
Barbula unguiculata Hedw.AAl: Sérgio et al., 1997b.
Ag: Sérgio et al., 1984.
BAl: Pereira-Coutinho, 1917b.
BL: Henriques, 1889; Machado, 1928; Sá-Nogueira, 1950a; Allorge, 1974; Sérgio et
al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1928; Allorge, 1974.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Machado, 1928; Barros,
1942b; Barros, 1942c; Barros, 1943; Allorge, 1974; Sérgio & Correia, 1981; Sérgio
& Sim-Sim, 1985a.
Mi: Machado, 1913; Machado, 1920a; Machado, 1928; Allorge, 1974; Sérgio &
Schumacker, 1992.
R: Sérgio et al., 1988.		Barbula unguiculata Barbula unguiculata
47
Bartramia ithyphylla Brid.IPmap: Casas et al., 1992.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Machado,
1930; Casas et al., 1992.
BA: Henriques, 1883, Henriques, 1889; Sérgio, 1969; Casas et al., 1992; Sérgio,
1994; Jansen 1998.
BA/BB: Levier, 1880; Pereira-Coutinho, 1917b; Machado, 1930; Greven & Melick,
1994.
Mi: Pereira-Coutinho, 1917b; Sá-Nogueira, 1950d; Casas et al., 1992; Sérgio &
Schumacker, 1992.		Bartramia ithyphylla Bartramia ithyphylla
48
Bartramia pomiformis Hedw.IPmap: Casas et al., 1992.
AAl: Allorge, 1974; Casas et al., 1992; Sérgio et al., 1997b.
Ag: Henriques, 1889; Dixon, 1912; Pereira-Coutinho, 1917b; Allorge, 1931a;
Allorge, 1974; Casas et al., 1992.
BA: Henriques, 1883; Henriques, 1889; Pereira-Coutinho, 1917b; Barros, 1942b;
Braun-Blanquet et al., 1956; Tavares, 1956; Allorge, 1974; Casas et al., 1992;
Sérgio, 1994a; Sérgio et al., 2001d.
BA/BB: Machado, 1917b; Pereira-Coutinho, 1917b; Greven & Melick, 1994.
BB: Luisier, 1916b; Casas et al., 1992; Jansen , 1994; Sérgio et al., 2002c.
BL: Brotero, 1804; Henriques, 1889; Pereira-Coutinho, 1917b; Sá-Nogueira, 1950a;
Allorge, 1974; Casas et al., 1992; Werner, 1997.
DL: Anónimo, 1883a; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Pereira-Coutinho, 1917b; Braun-Blanquet et al., 1964; Casas et al., 1992; Werner,
1997.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Casas et al., 1992.
Mi: Henriques, 1889; Anónimo, 1899; Sampaio, 1901-1902; Machado, 1913;
Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1920a; Barros, 1944b; Sá-
Nogueira, 1950d; Allorge, 1974; Casas et al., 1992; Sérgio & Schumacker, 1992.
R: Barros, 1942a; Allorge, 1974; Casas et al., 1992.
TM: Ervideira, 1926; Allorge & Allorge, 1949; Allorge, 1974; Casas et al., 1992;
Sérgio & Schumacker, 1992.
Pt: Ervideira, 1922; Machado, 1930.		Bartramia pomiformis Bartramia pomiformis
49
Bartramia stricta Brid.IPmap: Casas et al., 1992.
AAl: Luisier, 1916b; Allorge, 1974; Casas et al., 1992; Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Luisier, 1916b;
Sérgio, 1970-1971; Allorge, 1974; Guerra et al., 1981; Sérgio et al., 1984; Casas et
al., 1992.
BA: Braun-Blanquet et al., 1952; Sérgio, 1994a.
BAl: Sérgio, 1970-1971; Allorge, 1974; Casas et al., 1992; Sérgio et al.,1999a.
BB: Catarino et al., 1987; Casas et al., 1992.
BL: Henriques, 1889; Warnstorf, 1899; Pereira-Coutinho, 1917b; Allorge, 1974;
Catarino et al., 1987; Sérgio et al., 1988; Casas et al., 1992.
DL: Henriques, 1889; Sampaio, 1901-1902; Casas et al., 1992.
E: Mitten, 1853; Henriques, 1889; Luisier, 1916b; Pereira-Coutinho, 1917b; Barros,
1942b; Allorge, 1974; Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985a; Sérgio et
al., 1988; Casas et al., 1992.
Mi: Machado, 1913; Machado, 1920a; Sá-Nogueira, 1950c; Sá-Nogueira, 1950d;
Casas et al., 1992; Sérgio & Schumacker, 1992.
R: Pereira-Coutinho, 1917b; Barros, 1942a; Barros, 1942b; Allorge, 1974; Sérgio et
al., 1988; Casas et al., 1992.
TM: Henriques, 1889; Sampaio, 1901-1902; Machado, 1915b; Ervideira, 1922;
Barros, 1942b; Allorge & Allorge, 1949; Braun-Blanquet et al., 1956; Allorge, 1974;
Casas et al., 1992.
Pt: Luisier, 1924; Machado, 1930.		Bartramia stricta Bartramia stricta
50
Brachydontium trichodes (F. Weber) MildeDL: Machado, 1917d; Machado, 1928; Casares, 1932; Sérgio et al., 2001e.
It has not been seen since 1917, in spite of the careful searches performed in the area, so is
certainly extinct (Sérgio et al., 2001e).		Brachydontium trichodes Brachydontium trichodes
51
Brachythecium albicans (Hedw.) Schimp.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Machado, 1933; Sérgio et al., 2001e.
BA: Machado, 1917b; Sérgio, 1972a; Sérgio, 1994a; Jansen & Sequeira, 1999.
BA/BB: Greven & Melick, 1994.
BB: Luisier, 1916b; Machado, 1933; Sérgio et al., 2002c.
DL: Machado, 1933; Luisier, 1948.
Mi: Machado, 1914a; Pereira-Coutinho, 1917b; Machado, 1920a; Machado, 1933;
Barros, 1944b; Sá-Nogueira, 1950d.
TM: Machado, 1933; Allorge & Allorge, 1949; Sérgio & Schumacker, 1992.
See note under Brachythecium oxycladum.		Brachythecium albicans Brachythecium albicans
52
Brachythecium campestre (Müll. Hal.) Schimp.DL: Henriques, 1889; Machado, 1933.
Mi: Sá-Nogueira, 1950d; Sérgio, 1972a; Sérgio & Schumacker, 1992.
We believe that new localities can be found.		Brachythecium campestre Brachythecium campestre
53
Brachythecium dieckii RoellBA: Henriques, 1889; Machado, 1933; Sérgio, 1994a; Jansen 1998; Sérgio & Jansen,
2000.
BB: Machado, 1933.
TM: Sérgio & Jansen 2000.
Pt: Kindberg, 1887.
Based on the taxonomic revision of the B. velutinum group (Sérgio & Jansen, 2000), some
plants formerly considered as var. salicinum (Schimp.) Mönk. or B. venustum De Not.
correspond to this species, as it happen to the material of Beira Alta, Serra da Estrela, S.
Romão, leg. Levier (G!) first reported by Henriques (1889) as B. venustum.		Brachythecium dieckii Brachythecium dieckii
54
Brachythecium glareosum (Spruce) Schimp.BA: Pierrot, 1972 (Comm.), Sérgio, 1994a.
E: Machado, 1917a; Pereira-Coutinho, 1917b; Machado, 1933; Barros, 1943.
TM: Sérgio & Schumacker, 1992.		Brachythecium glareosum Brachythecium glareosum
55
Brachythecium mildeanum (Schimp.) MildeBA: Teles, 1970.
BB: Sérgio et al., 2002c.
E: Sérgio et al., 1987b; Sérgio et al., 1988.
TM: Teles, 1970; Sérgio & Schumacker, 1992.		Brachythecium mildeanum Brachythecium mildeanum
56
Brachythecium oxycladum (Brid.) A. Jaeger[Ag: Solms-Laubach, 1868; Henriques, 1889; Machado, 1933].
We have had the opportunity to study the only specimen recorded in Portugal (Solms-
Laubach, 1868) sub B. lactum (BM!). But in fact this material corresponds to
Brachythecium albicans.		Brachythecium oxycladum Brachythecium oxycladum
57
Brachythecium plumosum (Hedw.) Schimp.AAl/BAl: Machado, 1933.
BA: Sérgio, 1972a.
BA/BB: Greven & Melick, 1994.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1933.
Mi: Machado, 1913; Machado, 1916; Machado, 1920a; Machado, 1933; Barros,
1944b; Sá-Nogueira, 1950d; Sérgio & Schumacker, 1992.
TM: Machado, 1933.
Pt: Brotero, 1804.		Brachythecium plumosum Brachythecium plumosum
58
Brachythecium populeum (Hedw.) Schimp.BA: Machado, 1917a; Sérgio, 1994a.
BB: Machado, 1914a; Machado, 1916; Machado, 1917b; Machado, 1933; Allorge,
1974.
BL: Allorge, 1974.
Mi: Machado, 1933.		Brachythecium populeum Brachythecium populeum
59
Brachythecium rivulare Schimp.AAl: Sérgio et al., 1997b.
Ag: Pereira-Coutinho, 1917b; Machado, 1933.
BA: Pereira-Coutinho, 1917b; Machado, 1933; Sérgio, 1972a; Sérgio, 1994a.
BA/BB: Greven & Melick, 1994.
BAl: Sérgio et al.,1999a; Sérgio & Garcia, 2001b.
DL: Luisier, 1916b.
Mi: Henriques, 1889; Machado, 1913; Luisier, 1916b; Machado, 1917a; Machado,
1917b; Pereira-Coutinho, 1917b; Machado, 1920a; Machado, 1933; Barros, 1944b;
Sá-Nogueira, 1950d; Teles, 1970; Sérgio & Schumacker, 1992.
R: Sérgio et al., 1988.
TM: Ervideira, 1922; Allorge & Allorge, 1949; Teles, 1970.		Brachythecium rivulare Brachythecium rivulare
60
Brachythecium rutabulum (Hedw.) Schimp.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Allorge, 1974.
BA: Sérgio et al., 2001d.
BAl: Luisier, 1916b.
BL: Sá-Nogueira, 1950a; Sérgio, 1972a; Allorge, 1974; Sérgio et al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1933; Allorge, 1974.
E: Mitten, 1853; Henriques, 1889; Warnstorf, 1899; Pereira-Coutinho, 1917b;
Machado, 1933; Barros, 1943; Braun-Blanquet et al., 1964; Allorge, 1974; Sérgio &
Sim-Sim, 1985a.
Mi: Henriques, 1889; Machado, 1913; Machado, 1914a; Machado, 1916; Pereira-
Coutinho, 1917b; Machado, 1920a; Machado, 1933; Barros, 1944b; Sá-Nogueira,
1950d; Allorge, 1974; Sérgio & Schumacker, 1992.
R: Pereira-Coutinho, 1917b; Machado, 1933; Allorge, 1974.
TM: Allorge, 1974.
Some reports of B. rutabulum var. turgescens Limpr. are included in this species. The
mentioned variety was reported from different localities by Allorge (1974).		Brachythecium rutabulum Brachythecium rutabulum
61
Brachythecium salebrosum (F. Weber & D. Mohr) Schimp.AAl/BAl: Henriques, 1889; Machado, 1933; Allorge, 1974.
Ag: Solms-Laubach, 1868; Henriques, 1889; Machado, 1933; Allorge, 1974.
BA: Sérgio et al., 2001d.
BAl: Sérgio, 1970-1971; Allorge, 1974.
BL: Allorge, 1974.
E: Barros, 1943; Barros, 1944a; Allorge, 1974.
Mi: Allorge, 1974.		Brachythecium salebrosum Brachythecium salebrosum
62
Brachythecium starkei (Brid.) Schimp.BA: Machado, 1917a; Machado, 1917b; Machado, 1933; Sérgio, 1994a; Sérgio et
al., 2001b; Sérgio et al., 2002a.
DL: Henriques, 1889; Machado, 1933.
Pt: Luisier, 1924.
Its occurrence in Beira Alta and Portugal is confirmed on the basis of new collections. So,
now it is to be considered a species not extinct from Portugal (Sérgio et al., 2002a).		Brachythecium starkei Brachythecium starkei
63
Brachythecium velutinum (Hedw.) Schimp.AAl: Sérgio et al., 1997b.
Ag: Sérgio, 1970-1971; Allorge, 1974.
BA: Henriques, 1889; Machado, 1917b; Machado, 1933; Allorge, 1974; Sérgio,
1994a.
BA/BB: Sampaio, 1901-1902; Greven & Melick, 1994.
BAl: , Sérgio et al., 2002d.
BL: Warnstorf, 1899; Allorge, 1974; Sérgio & Sim-Sim, 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Machado, 1933; Allorge, 1974.
E: Machado, 1933; Allorge, 1974; Sérgio, 1981; Bento-Pereira & Sérgio, 1983;
Sérgio & Sim-Sim, 1985d.
Mi: Machado, 1914a; Machado, 1915a; Pereira-Coutinho, 1917b; Machado, 1920a;
Machado, 1933; Allorge, 1974.
R: Sérgio et al., 1998d.
TM: Ervideira, 1922; Machado, 1933; Allorge, 1974; Sérgio & Schumacker, 1992.		Brachythecium velutinum Brachythecium velutinum
64
Bruchia vogesiaca Schwägr.Eurmap: Bonnot, 1967b; Sérgio et al., 1998b.
IPmap: Elias, 1987-1988; Sérgio et al., 1994c., Casas et al., 1996; Sérgio & Draper,
2002a; Sérgio & Draper, 2002b.
BA: Séneca & Sérgio, 1992; Sérgio, 1994a; Casas et al., 1996; Sérgio et al., 1998b.
BB: Luisier, 1913; Machado, 1928; Casares, 1932; Elias, 1987-1988; Bonnot,
1967b; Casas et al., 1996; Sérgio et al., 1998b.
Mi: Machado, 1918a; Machado, 1920a; Machado, 1928; Casares, 1932; Bonnot,
1967b; Sérgio et al., 1998b; Rushing, 1986; Elias, 1987-1988; Séneca & Sérgio,
1992; Casas et al., 1996.
Pt: Bonnot, 1967a; Sérgio et al., 1994c.
For a review of this endangered species see Sérgio et al. (1998b).		Bruchia vogesiaca Bruchia vogesiaca
65
Bryoerythrophyllum campylocarpum (Müll. Hal.) Crum.IPmap: Sérgio et al., 1994c.
BL: Allorge, 1974.
DL: Luisier, 1948.
Mi: Machado, 1917a; Pereira-Coutinho, 1917b; Machado, 1920a; Machado, 1928;
Casares, 1932; Sérgio, 1968-1969; Hill, 1981; Sollman, 1990; Casas et al., 1996;
Sérgio, 1997b; Sérgio et al., 2001e.
Pt: Luisier, 1934; Sérgio, 1968-1969.
Firstly described by Machado (1917a, Orig. descrip.), for Minho (Paredes de Coura, Mantelães) sub Hyophila crenulata, was later synonymised to Hyophila machadoana Sérgio and Bryoerythrophyllum machadoanum (Sérgio) M. O. Hill (Sérgio, 1968-1969;
Hill, 1981). Later, Sollman (1990) affirmed that this species is the same taxa of B. campylocarpum. For other information on this species see under Amphidium lapponicum.		Bryoerythrophyllum campylocarpum Bryoerythrophyllum campylocarpum
66
Bryoerythrophyllum lusitanicum (Cardot & Dixon) M. Hill.BL: Dixon, 1912 (Orig. descrip.), G. Roth, 1913; Machado, 1928; Casares, 1932;
Allorge, 1974; Hill, 1981; Zander, 1986.
Pt: Luisier, 1934.
Dixon (1912) described this taxon as a new species of portuguese bryoflora. The
interpretation of the mentioned taxon, accordind to Soll		Bryoerythrophyllum lusitanicum Bryoerythrophyllum lusitanicum
67
Bryum alpinum With. s. l.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Dixon, 1912;
Pereira-Coutinho, 1917b; Allorge, 1931a; Sérgio, 1970-1971; Allorge, 1974.
BA: Henriques, 1883, Henriques, 1889; Luisier, 1916b; Pereira-Coutinho, 1917b;
Pierrot, 1972; Allorge, 1974; Sérgio, 1994a; Jansen 1998; Jansen & Sequeira, 1999;
Sérgio et al., 2001d.
BA/BB: Levier, 1880; Machado, 1917b; Pereira-Coutinho, 1917b; Greven & Melick,
1994.
BB: Sérgio et al., 2002c.
BL: Allorge, 1935; Allorge, 1974; Sérgio et al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Werner, 1997.
E: Henriques, 1889; Sérgio & Tauleigne Gomes, 2001.
Mi: Anónimo, 1887; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Machado, 1913; Machado, 1914a; Machado, 1917b; Pereira-Coutinho, 1917b;
Machado, 1920a; Barros, 1944b; Sá-Nogueira, 1950d; Allorge, 1974; Sérgio &
Schumacker, 1992.
R: Sérgio et al., 1988.
TM: Machado, 1915b; Ervideira, 1922; Ervideira, 1926; Allorge, 1974; Sérgio &
Schumacker, 1992.
Pt: Luisier, 1924; Machado, 1930.
Many reports of this species belong to var. meridionale Schimp., that is in fact the more
frequent variety in Portugal.		Bryum alpinum Bryum alpinum
68
Bryum argenteum Hedw.AAl: Barros, 1942b; Allorge, 1974; Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Allorge,
1931a; Sérgio et al., 1984.
BA: Sérgio, 1994a.
BA/BB: Greven & Melick, 1994.
BB: Machado, 1917b; Séneca-Cardoso, 1989.
BL: Brotero, 1804; Henriques, 1889; Pereira-Coutinho, 1917b; Allorge, 1931a;
Allorge, 1974; Sérgio et al., 1988.
DL: Anónimo, 1883a; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Pereira-Coutinho, 1917b.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Barros, 1942b; Barros,
1942c; Allorge, 1974; Sérgio, 1981; Sérgio & Correia, 1981; Bento-Pereira &
Sérgio, 1983; Sérgio & Sim-Sim, 1985a; Sérgio & Sim-Sim, 1985d; Sérgio, 1987b.
Mi: Sampaio, 1901-1902; Machado, 1913; Luisier, 1916b; Machado, 1920a;
Allorge, 1974; Sérgio & Schumacker, 1992.
R: Sérgio et al., 1988.
TM: Pereira-Coutinho, 1917b; Ervideira, 1922; Ervideira, 1926; Sérgio &
Schumacker, 1992.
Pt: Machado, 1930.		Bryum argenteum Bryum argenteum
69
Bryum caespiticium Hedw.AAl: Sérgio et al., 1997b.
Ag: Machado, 1930; Sérgio et al., 1984.
BA: Machado, 1917a; Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1918b;
Machado, 1930; Sérgio, 1969; Teles, 1970; Sérgio, 1994a.
BA/BB: Machado, 1930.
BL: Brotero, 1804; Sérgio, 1969; Sérgio et al., 1988.
E: Ade & Koppe, 1942; Sérgio, 1970-1971; Pierrot, 1972; Sérgio & Correia, 1981;
Sérgio & Sim-Sim, 1985a; Sérgio & Sim-Sim, 1985d; Sérgio et al., 1988.
Mi: Luisier, 1907a; Machado, 1913; Machado, 1917a; Machado, 1930.
R: Sérgio et al., 1988.
TM: Machado, 1915b; Pereira-Coutinho, 1917b; Machado, 1918b; Ervideira, 1926;
Machado, 1930; Teles, 1970.
Pt: Luisier, 1924.
We have included in the type variety the reports of Bryum caespiticium var. pseudokunzei
(Limpr.) A. Mach. and Bryum caespiticium Hedw. var. comense (Schimp.) A. Mach.
Without new studies these varieties can not be considered taxonomically significant.		Bryum caespiticium Bryum caespiticium
70
Bryum canariense Brid.Ag: Solms-Laubach, 1868; Welwitsch, 1872; Henriques, 1889; Dixon, 1912;
Pereira-Coutinho, 1917b; Machado, 1930; Sérgio, 1970-1971.
BA: Sérgio, 1969.
BL: Sérgio et al., 1988.
E: Pierrot, 1972; Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985a.
TM: Allorge, 1935; Allorge & Allorge, 1949.
Pt: Luisier, 1924.
The var. provinciale (Philib.) Husn. is not separeted from the var. type by several authors,
however the two varieties are included in the check-list of the mosses of Italy by Cortinini
Pedrottti (2001). Only reported from Beira Alta in one locality by Sérgio (1969). Future
observations could confirm the taxonomic status of this taxa in Portugal.		Bryum canariense Bryum canariense
71
Bryum capillare Hedw.AAl: Catarino et al., 1987; Sérgio et al., 1997b; Sérgio et al., 1998d.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Pereira-
Coutinho, 1917b; Sérgio, 1970-1971; Sérgio et al., 1984.
BA: Henriques, 1883; Henriques, 1889; Sérgio, 1994a; Jansen 1998; Sérgio et al.,
2001d.
BA/BB: Levier, 1880; Greven & Melick, 1994.
BAl: Sérgio, 1970-1971; Sérgio et al.,1999a; Sérgio et al., 2002d.
BB: Luisier, 1916b; Catarino et al., 1987; Sérgio et al., 2002c.
BL: Brotero, 1804; Henriques, 1889; Barros, 1942a; Sérgio et al., 1988; Sérgio &
Sim-Sim, 1988.
DL: Henriques, 1889; Sampaio, 1901-1902.
E: Mitten, 1853; Henriques, 1889; Luisier, 1916b; Pereira-Coutinho, 1917b; Barros,
1942a; Barros, 1942b; Barros, 1942c; Barros, 1943; Sá-Nogueira, 1950b; Sérgio,
1970-1971; Sérgio, 1981; Sérgio & Correia, 1981; Bento-Pereira & Sérgio, 1983;
Sérgio & Sim-Sim, 1985a; Sérgio & Sim-Sim, 1985d; Sérgio et al., 1988.
Mi: Machado, 1913; Machado, 1920a; Barros, 1944b; Sá-Nogueira, 1950d; Sérgio &
Schumacker, 1992.
R: Barros, 1942a; Catarino et al., 1987; Sérgio et al., 1988; Sérgio, C. & Sim-Sim,
M., 1994; Sérgio et al., 1998d.
TM: Ervideira, 1926; Sérgio & Schumacker, 1992.
Pt: Ervideira, 1922; Machado, 1930.
We have included in the type variety the reports of B. capillare var. meridionale Schimp.		Bryum capillare Bryum capillare
72
Bryum dichotumum Hedw.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Welwitsch, 1872; Henriques,
1889; Dixon, 1912; Pereira-Coutinho, 1917b; Allorge, 1931a; Sérgio, 1970-1971;
Allorge, 1974; Sérgio et al., 1984.
BA/BB: Greven & Melick, 1994.
BAl: Sérgio, 1970-1971; Allorge, 1974.
BB: Sérgio et al., 2002c.
BL: Levier, 1880; Henriques, 1889; Pereira-Coutinho, 1917b; Allorge, 1931a (Type
material ? of var. pseudo-blindii Amann); Sá-Nogueira, 1950a; Allorge & Casas de
Puig 1958; Allorge, 1974; Sérgio et al., 1988; Sérgio & Sim-Sim, 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Séneca-Cardoso, 1989.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Ade & Koppe, 1942;
Barros, 1942a; Barros, 1942b; Barros, 1943; Sá-Nogueira, 1950b; Allorge, 1974;
Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985d; Sérgio et al., 1988.
Mi: Henriques, 1889; Sampaio, 1901-1902; Machado, 1913; Machado, 1920a;
Barros, 1944b; Sá-Nogueira, 1950d; Allorge, 1974; Sérgio & Schumacker, 1992.
R: Pereira-Coutinho, 1917b; Barros, 1942a; Barros, 1942b; Allorge, 1974; Sérgio et
al., 1988.
TM: Machado, 1915b; Ervideira, 1922.
Pt: Machado, 1930.
This is a polymorphic taxon (= Bryum bicolor Dicks.) that was considered complex that
includes different species as Bryum barnesii J. B. Wood. However this species is now
indicated as a distinct species (Vanderpoorten & Zartman, 2002). So, the distribution of
this species may change a lot after the revision of the different herbarium specimens.		Bryum dichotumum Bryum dichotumum
73
Bryum donianum Grev.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Dixon, 1912; Luisier, 1916b; Pereira-
Coutinho, 1917b; Allorge, 1931a; Allorge, 1974; Sérgio et al., 1984.
BA: Sérgio et al., 2001d.
BAl: Sérgio & Garcia, 2001b.
BL: Henriques, 1889; Dixon, 1912; Pereira-Coutinho, 1917b; Allorge, 1974; Sérgio
et al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Séneca-Cardoso, 1989.
E: Henriques, 1889; Warnstorf, 1899; Pereira-Coutinho, 1917b; Ade & Koppe,
1942; Barros, 1942a; Allorge, 1974; Sérgio & Sim-Sim, 1985a.
Mi: Pereira-Coutinho, 1917b; Machado, 1920a; Sá-Nogueira, 1950d; Allorge, 1974;
Sérgio & Schumacker, 1992.
Pt: Venturi, 1881; Luisier, 1924; Machado, 1930.		Bryum donianum Bryum donianum
74
Bryum dunense A. J. E. Sm. & H. WhitehouseAg: Sérgio et al., 1984.
BL: Leitão et al., 1996.
E: Guerra & Puche, 1984.
Mi: Guerra & Puche, 1984.
Based on the taxonomic study of Bryum bicolor complex (Vanderpoorten & Zartman,
2002) this species is considered a synonym of B. bicolor see B. dichotumum). For
Portugal, all taxa of this complex (B. bicolor, B. gemmiluscens, B. gemmiferum and B.
barnesii) need to be revised.		Bryum dunense Bryum dunense
75
Bryum elegans NeesAg: Dixon, 1912; Machado, 1930.
BA: Sérgio, 1994a.
BA/BB: Pereira-Coutinho, 1917b; Machado, 1930; Greven & Melick, 1994.
Mi: Machado, 1920a.
Pt: Luisier, 1924; Machado, 1930.		Bryum elegans Bryum elegans
76
Bryum gemmiferum R. Wilczek & DemaretAg: Sérgio et al., 1984.
BL: Sérgio et al., 1988.
R: Sérgio et al., 1988.		Bryum gemmiferum Bryum gemmiferum
77
Bryum gemmilucens R. Wilczek & DemaretIPmap: Elias, 1988.
Ag: Sérgio et al., 1984.
BB: Sérgio et al., 2002c.
BL: Sérgio et al., 1988.		Bryum gemmilucens Bryum gemmilucens
78
Bryum gemmiparum De Not.AAl: Sérgio et al., 1997b.
Ag: Dixon, 1912; Machado, 1930; Allorge, 1931a; Allorge, 1974; Sérgio et al.,
1984.
BL: Allorge, 1931a; Allorge, 1974.
E: Allorge, 1931a; Allorge, 1974; Sérgio, 1981; Sérgio & Correia, 1981; Bento-
Pereira & Sérgio, 1983; Sérgio & Sim-Sim, 1985a; Sérgio & Sim-Sim, 1985d.		Bryum gemmiparum Bryum gemmiparum
79
Bryum klinggraeffii Schimp.Ag: Townsend, 1994.Bryum klinggraeffii Bryum klinggraeffii
80
Bryum machadoi G. Roth.A species erroneously reported from Portugal by Podpera, 1954. Moenkemeyer (1927)
reported the species to �Salamanka in Portugal� based on a collection from Machado in
1914 as Bryum pendulum subsp. machadoi (G. Roth) Amann. This report is to be placed
in Spain.		Bryum machadoi Bryum machadoi
81
Bryum mildeanum Jur.Ag: Sérgio et al., 1984.
BA: Teles, 1970.
E: Sérgio et al., 1987b.
TM: Teles, 1970.		Bryum mildeanum Bryum mildeanum
82
Bryum minii Podp.Ipmap: Sérgio & Draper, 2002b.
Ptmap: Sérgio et al., 1999b.
AAl: Pierrot, 1972; Sérgio et al., 1997b; Sérgio et al., 1999b.
BA: Machado, 1918; Machado, 1930; Machado, 1935; Jansen & Sequeira, 1999;
Sérgio et al., 1999b; Sérgio et al., 2001d.
BB: Sérgio, 1968-1969; Sérgio et al., 1999b; Sérgio et al., 2001d; Sérgio et al.,
2001c; Sérgio et al., 2002c.
DL: Pereira-Coutinho, 1917b; Sérgio et al., 1999b.
Mi: Machado, 1916; Machado, 1918; Machado, 1920a; Machado, 1930; Machado,
1935; Sérgio & Schumacker, 1992; Sérgio et al., 1999b.
Pt: Luisier, 1924; Podpera, 1954; Podpera, 1955; Podpera, 1973.
The first published record of this species in Portugal was provided by Machado (1916)
who cited a specimen of Bryum marginatum Schimp. collected in Minho, Famalicão, in
the northern part of the country. Machado (1935) considered that this species should be
included in the Pseudotriquetra Section as a new species, B. minii Podp. in litt. based on
Podepera observations (Orig. descrip.). Sérgio (1968-1969) reported this species again at
a new locality, Beira Baixa, and more recently it was considered a more or less frequent
species in Portugal (Sérgio et al., 1999b).		Bryum minii Bryum minii
83
Bryum muehlenbeckii Bruch & Schimp.AAl: Sérgio et al., 1997b.
BA: Machado, 1920b; Machado, 1930; Sérgio, 1994a.
Mi: Sérgio & Schumacker, 1992.
R: Ferreira, 1992.
Pt: Machado, 1930.		Bryum muehlenbeckii Bryum muehlenbeckii
84
Bryum neodamense Itzig. ex Müll. Hal.AAl: Sérgio & Casas, 2002.Bryum neodamense Bryum neodamense
85
Bryum pallescens Schwägr.BA: Machado, 1917b; Machado, 1933; Sérgio, 1968-1969; Sérgio, 1994a.
BB: Pereira-Coutinho, 1917b.
TM: Allorge & Allorge, 1949.		Bryum pallescens Bryum pallescens
86
Bryum platyloma Schwägr.Mi: Machado, 1917a; Machado, 1917b; Machado, 1930; Sá-Nogueira, 1950d; Sérgio
& Schumacker, 1992.
Pt: Luisier, 1934.		Bryum platyloma Bryum platyloma
87
Bryum pseudotriquetrum (Hedw.) P. Gaertn., B. Mey. & Scherb.AAl: Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Henriques, 1889; Dixon, 1912; Pereira-Coutinho, 1917b;
Machado, 1930; Guerra et al., 1981.
BA: Teles, 1970; Pierrot, 1972; Jansen & Sequeira, 1999; Sérgio et al., 2001d.
BA/BB: Greven & Melick, 1994.
BB: Sérgio et al., 2002c.
BAl: Sérgio et al.,1999a; Sérgio & Garcia, 2001b.
BL: Allorge, 1931a; Sá-Nogueira, 1950a; Werner, 1997.
DL: Henriques, 1889; Sampaio, 1901-1902.
E: Sá-Nogueira, 1950e; Sérgio et al., 1988.
Mi: Machado, 1914a; Machado, 1917b; Pereira-Coutinho, 1917b; Machado, 1920a;
Machado, 1930; Barros, 1944b; Sá-Nogueira, 1950d; Teles, 1970; Sérgio &
Schumacker, 1992.
TM: Ervideira, 1922; Allorge & Allorge, 1949; Teles, 1970; Séneca-Cardoso, 1989.
Pt: Machado, 1930.		Bryum pseudotriquetrum Bryum pseudotriquetrum
88
Bryum radiculosum Brid.AAl: Sérgio et al., 1997b.
Ag: Dixon, 1912; Machado, 1930.
BL: Warnstorf, 1899.
DL: Anónimo, 1883a; Henriques, 1889; Anónimo, 1890; Sampaio, 1901-1902;
Luisier, 1916b; Pereira-Coutinho, 1917b.
E: Barros, 1943; Sérgio & Tauleigne Gomes, 2001.
Mi: Machado, 1920a; Barros, 1944b; Sá-Nogueira, 1950d; Sérgio & Schumacker, 1992.		Bryum radiculosum Bryum radiculosum
89
Bryum rubens Mitt.Ag: Townsend, 1994.
BA: Sérgio et al., 2001a.
BL: Allorge, 1974; Sérgio, 2002b.
E: Pereira-Coutinho, 1917b; Machado, 1930; Allorge, 1974.
Mi: Machado, 1914a.		Bryum rubens Bryum rubens
90
Bryum ruderale Crundw. & NyholmTM: Pierrot, 1972.Bryum ruderale Bryum ruderale
91
Bryum schleicheri Schwägr.BA: Sérgio, 1968-1969.Bryum schleicheri Bryum schleicheri
92
Bryum subapiculatum HampeBA: Sérgio et al., 2001d.Bryum subapiculatum Bryum subapiculatum
93
Bryum torquescens De Not.AAl: Pereira-Coutinho, 1917b; Sérgio et al., 1997b.
Ag: Solms-Laubach, 1868; Estácio da Veiga, 1869; Henriques, 1889; Dixon, 1912;
Sérgio, 1970-1971; Allorge, 1974; Sérgio et al., 1984.
BA: Sérgio, 1994a.
BAl: Pereira-Coutinho, 1917b; Sérgio, 1970-1971; Allorge, 1974; Sérgio et
al.,1999a.
BL: Henriques, 1889; Pereira-Coutinho, 1917b; Allorge, 1974; Sérgio et al., 1988.
DL: Henriques, 1889; Sampaio, 1901-1902; Pereira-Coutinho, 1917b.
E: Mitten, 1853; Henriques, 1889; Pereira-Coutinho, 1917b; Sérgio, 1970-1971;
Allorge, 1974; Sérgio & Correia, 1981; Sérgio & Sim-Sim, 1985a; Sérgio et al.,
1988.
Mi: Sampaio, 1901-1902; Machado, 1913; Machado, 1920a.
R: Pereira-Coutinho, 1917b; Sérgio et al., 1988.
TM: Machado, 1915b.
Pt: Machado, 1930.		Bryum torquescens Bryum torquescens
94
Bryum valparaisense Thér.Wmap: Arts et al., 1995.
Ag: Townsend, 1994.
BAl: Sérgio & Garcia, 2001b.		Bryum valparaisense Bryum valparaisense
95
Buckiella undulata (Hedw.) IrelandIPmap: Casas et al., 1992.
[E: Henriques, 1889; Machado, 1933].
Mi: Machado, 1920a; Machado, 1933; Casas et al., 1992; Sérgio & Schumaker,
1992.
TM: Casas et al., 1992.
The species was reported dubiously to Estremadura, Mafra. In fact, all the reports
gathered by E. da Veiga and reported by Henriques or Machado are uncertain.		Buckiella undulata Buckiella undulata
96
Calliergonella cuspidata (Hedw.) LoeskeAg: Machado, 1933; Allorge, 1974.
BA: Luisier, 1907a; Luisier, 1916b; Pereira-Coutinho, 1917b; Sérgio, 1969; Teles,
1970; Allorge, 1974; Sérgio, 1994a.
BB: Machado, 1933.
BL: Allorge, 1931a; Machado, 1933; Anónimo, 1969; Allorge, 1974.
DL: Sampaio, 1901-1902; Machado, 1933; Sá-Nogueira, 1958; Allorge, 1974.
E: Luisier, 1910c; Luisier, 1916b; Machado, 1933; Allorge, 1974.
Mi: Henriques, 1889; Machado, 1913; Luisier, 1916b; Pereira-Coutinho, 1917b;
Machado, 1920a; Machado, 1933; Sá-Nogueira, 1950d; Teles, 1970; Allorge, 1974;
Sérgio & Schumacker, 1992.
TM: Ervideira, 1922; Machado, 1933; Allorge & Allorge, 1949; Teles, 1970;
Allorge, 1974; Sérgio & Schumacker, 1992.		Calliergonella cuspidata Calliergonella cuspidata
97
Campyliadelphus chrysophyllus (Brid.) R. S. ChopraBL: Allorge, 1931b; Machado, 1933; Allorge, 1974; Sérgio et al., 2001e.
Is certainly an extinct species (Sérgio et al., 2001e).		Campyliadelphus chrysophyllus Campyliadelphus chrysophyllus
98
Campylium stellatum (Hedw.) C. E. O. JensenBL: Sérgio, 1968-1969; Anónimo, 1969.
Only reported from Vouga region, Pateira de Fermentelos. The two specimens in
COI were recently confirmed (!).		Campylium stellatum Campylium stellatum
99
Campylopus atrovirens De Not.[E: Machado, 1928].
The report of its occurrence in Portugal, Estremadura: Arrentela and Algarve, is still
uncertain, as Machado (1928) could not validate the species identification (nom vidi). In
LISU the specimen from Arrentela (Leg. Welwitsch) was not found.		Campylopus atrovirens Campylopus atrovirens
100
Campylopus brevipilus Bruch & Schimp.AAl: Sérgio et al., 1997b.
Ag: Estácio da Veiga, 1869; Henriques, 1889; Dixon, 1912; Machado, 1928;
Casares, 1932; Sérgio, 1970-1971; Allorge, 1974.
BA/BB: Greven & Melick, 1994.
BB: Braun-Blanquet et al., 1964.
BL: Allorge, 1935; Anónimo, 1967; Allorge, 1974.
DL: Werner, 1997.
E: Pereira-Coutinho, 1917b; Machado, 1928; Casares, 1932; Sérgio, 1970-1971;
Allorge, 1974; Sérgio & Sim-Sim, 1985a.
Mi: Pereira-Coutinho, 1917b; Machado, 1928; Allorge, 1931a; Casares, 1932; Sá-
Nogueira, 1950c; Sá-Nogueira, 1950d; Braun-Blanquet et al., 1964; Allorge, 1974.
R: Sérgio et al., 1988.
TM: Ervideira, 1922; Sérgio & Schumacker, 1992.
Campylopus brevipilus Campylopus brevipilus