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Introduction to plant parts

Diploma in Pharmacy

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Organization of plant cell

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Cell

Cells are the units of structure and functions in the bodies of all living organisms.

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  • Plant cell has three distinct regions
  • Cell wall
  • Protoplasm
  • Vacuole

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Cell organelles

In cell biology, an organelle is a specialized subunit usually within a cell that has a specific function.

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Cell inclusions (ergastic substances)

  • The non living substances of plant metabolism are termed as ergastic substances. These may be reserve foods, excretory products and secretory products of metabolism.

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  • Types of ergastic substances
    • Reserve foods
    • Excretory products
    • Secretory products

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1. Reserve foods

    • Carbohydrates
      • Many types of water soluble or water insoluble carbohydrates may be present. Glucose, fructose and sucrose are all water soluble.
      • Inulin is the only polysaccharide soluble in water which form sphere crystals.
      • Starch is another polysaccharide which is found in forms like granules.
      • Cellulose is a complex polysaccharide forming cell wall.
      • Mucilage which forms gummy solution is another type of polysaccharide.
    • Proteins
      • These are nitrogenous substances either soluble or insoluble in water. They may be amorphous (eg aleuronic layer) or crystalline (aleuronic grain) .They are converted to amino acids by proteolytic enzymes.
    • Lipids
      • These are reserve food materials of plants. Those which are liquid at room temperature are called oils and solids are termed as fats.

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2. Excretory products

    • Alkaloids
      • Generally speaking, alkaloids are those compounds that contain N in a heterocyclic ring. These are protective materials of plants discouraging animal or insect attacks. Alkaloids are also detoxifying agents of plants and reservoirs of protein synthesis.
    • Glycosides
      • Glycosides are condensation products of sugar and glycol.
    • Tannins
      • Tannins are astringent water soluble constituents present in the cell sap.
    • Resins
      • Resins water insoluble exudates derived from living sources- largely plants.
    • Latex
      • Latex is white suspension wherein microscopically small particles or oil globulesare suspended. Latex is present in lactiferous tissue characteristic to certain families like Euphorbiaceae, Papaveraceae etc
    • Volatile oil
      • These are water insoluble fragrant liquids found in the plant. They are also called essential oils. Normally volatile oils are carminative, stimulant and antiseptic.
    • Mineral crystals
      • These are water insoluble mineral aggregates which occur in cell wall or cell cavity. Common crystals found are calcium oxalate, calcium carbonate and silica etc

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3. Secretory products

      • These include enzymes, coloring substances/pigments and nectar

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Plant tissues

  • Plant tissue is a collection of similar cells performing an organized function for the plant.

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Classification of Plant Tissues

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Meristematic tissue

    • These cells have capacity to multiply. These are small isodiametric cells. These cells are compact, thinwalled, normally cubical and possess prominent nucleus.

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Simple permanent tissue

  • They are not usually changed to any other kind to tissue and normally retain their structural and physiological characters. These cells may be living or dead. These cells vary in shape, size and nature of protoplast.

  • Epidermis
    • It is the type of parenchymatous tissue forming the outermost layer of the plant. The outer layer of epidermal cells are often thick and covered with a fatty substance cutin. Its function is to conserve moisture and protect against mechanical injuries.

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  • Parenchyma
    • Parenchymatous cells are thinwalled polyhedral cells with large central vacuole. Aging results in change in shape, wall thickening and appearance of intercellular spaces.

  • Collenchyma
    • It is the supporting tissue found underneath the epidermis of many stems, leaf stalks and midribs. It is similar to parenchyma in structure except that primary cell wall is thickended to give mechanical strength of plant parts.

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  • Sclerenchyma
    • It is a supporting tissue and is purely mechanical in function. It is found in stems, roots, leaves and other parts along with xylem.

  • Cork
    • It is a simple permanent tissue made up of compactly arranged approximately prismatic cells found in radial rows. It forms the bark of stems or roots. Protoplasm in the cork cells dies hence matured cork cells are dead.

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Complex permanent tissue

  • This include xylem, phloem and secretory tissues
    • Xylem
      • It is the water conducting tissue of plant. The components of xylem are parenchyma, fibers and tracheary elements.Tracheary elements include tracheids and vessels.
    • Pholem
      • It is the food conducting tissue of the plant. The components of phloem are phloem sieve tubes, companion cells, phloem parenchyma and phloem fibers.
    • Secretory tissues
      • These are responsible for secretions which may be single celled or multicellular. They may be formed by splitting of cells ie schizogenous or by breakdown of cells ie lysigenous or by combnation of the former and later ie schizolysigenous. The secretions can be tannins, oils (volatile and fixed), mucilage and crystals.
      • Lactiferous tissue also known as milk tubes are the tissue belonging to some families that produce white juice known as latex.

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Xylem

Phloem

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Types of vascular bundles

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Morphology of Plant parts

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Root and Rhizome

  • Root: that portion of the plant axis which ascends into the soil and is devoid of the leaves
  • Rhizome: an underground stem which bears both roots and leaves eg Ginger

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  • The rhizomes are the stem structure which can be distinguished from roots by the presence of
    • Nodes and internodes
    • Scale leaves
    • Buds

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Morphology of Root and Rhizome

Shape

    • Cylindrical: circular in outline
    • Fusiform: thickened at the middle and tapering at the ends.
    • Conical: broad at the base and tapers towards the apex.
    • Napiform almost spherical with tapering at the lower part.
    • Tuberous: that contain tubers
    • Subcylindrical: nearly cylindrical

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  • Surface characters
    • Surface – smooth or rough
    • Presence or absence of scale leaves, nodes or internodes, scars of roots and leaves, transverse ridges, furrows.
    • Regular or irregular ridges.

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Fracture

    • Short: smooth fractured surface
    • Fibrous: fine threads extending from broken surface
    • Splintery: sharp and jagged projections on the fractured surface
    • Horny: Horn like broken surface
    • Laminated: Breaking into arranged layer
    • Granular: Fractured surface exhibiting small rounded prominences

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Leaves

  • Leaves are the lateral outgrowth of plant from stems.

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  • Morphology
  • Shape:
    • Acicular: needle shaped
    • Cordate: Heart shaped
    • Reniform: Kidney shaped
    • Lanceolate: lance shaped
    • Obovate: narrowing base and broading toward the apex
    • Oblong: long, wide with parallel margins
    • Ovate: broad at the base and tapering towards apex

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  • Apex
    • Cuspidate: having sharp end like point of the spear
    • Acute: ending in a sharp point
    • Obtuse: rounded or blunt end
    • Mucronate: terminating abruptly into a short and sharp point.
    • Enmarginate: having deep notch at the apex

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  • Base
    • Symmetrical: two sides of midrib being alike
    • Asymmetrical: two sides of midrib being unlike
    • Decurrent: leaf blade extends down the petiole
    • Cordate: heart shaped

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  • Margin
    • Entire: with continuous margin
    • Crenate: rounded teeth
    • Dentate: toothed and teeth directed outwardly
    • Serrate: having teeth of saw and teeth directed upwardly
    • Ciliate: with hairs

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  • Surface
    • Glabrous: smoothed surface or free from hairs and any other outgrowths
    • Rough: surface is somewhat harsh to tough
    • Glaucous: shiny green
    • Pubescent: covered with straight, short hairs
    • Gland dotted: Having non-sticky glands

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  • Type
    • Simple: These leaves cant be subdivided to leaflets.
    • Compound: that has number of leaflets.

# If as single, two, three, four or more leaflets are articulated to the petiole, it is called unifoliate, bifoliate, trifoliate, quadrafoliate, multifoliate etc

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Stomata

  • Stomata refers to a small pore on leaf which are meant for gaseous exchange and transpiration.
  • Stomata constitute of cell pore, guard cells and subsidiary cells.
  • # Transpiration is the process of water movement through a plant and its evaporation from aerial parts such as leaves, stem, and flowers.

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Types of stomata

  • Anomocytic type (irregular celled)
    • Stomata is surrounded by varying no of subsidiary cells Eg Digitalis
  • Anisocytic type (unequal celled)
    • Covered by three subsidiary cell and in that one is small in size than other two Eg Datura
  • Paracytic type (parallel celled)
    • 2 of the subsidiary cells have long axes parallel to the pore Eg Senna
  • Diacytic type (cross celled)
    • Guard cells are covered by two subsidiary cells on right angle to that of stomata. Eg Peppermint

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Trichome

  • An elongated tubular outgrowth of an epidermal cell is termed as trichome.

  • Functions of trichomes
    • Ion exchange
    • Protective
    • Secretion of essential oil
    • Absorption of water

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Types of trichome

  • Covering trichome
    • These are the trichomes that have apex and provide physical protection.
  • Glandular trichome
    • These are the trichomes that contain glandular head and secrete secondary metabolites.

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  • Covering trichomes can be unicellular, multicellular unbranched and multicellular branched.

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  • Glandular trichomes present in various plants

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Bark

  • All the secondary external tissues lying outside cork cambium is referred as bark.
  • Bark usually consist of cork (phellem), cork cambium (phellogen) and secondary cortex (phelloderm)

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  • Shape
    • Curved: concavity on the innerside
    • Recurved: concavity on the outer side
    • Channeled: concavity on the inner side is so great to form a deep channel
    • Quill: one edge overlapping other
    • Double quill: both edge rolling independently into two separate quills.
    • Compound quill: quill are packed one inside other.

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  • Characters of outer surface
    • Smooth or rough
    • Presence or absence of lenticels
    • Exfoliates, cracks, fissures, wrinkle, rhytidoma, epiphytes
  • Characters of inner surface
    • Striations (fine or coarse longitudinal ridges)
    • Corrugations (transverse wrinkles)

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Flower

  • It is the specialized shoot which is meant for the sexual reproduction of the plant.
  • A typical flower consist of calyx, corolla, androecium and gynoecium arranged into four separate whorls.

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  • Pedicel
    • Pedicilate (long pedicel)
    • Subsessile (a flower with a very short petiole)
    • Sessile (a flower with no pedicel)

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  • Type of flower
    • On the basis of sex organs
      • Unisexual flower flower having either androecium
      • Hermaphrodite: flower having both reproductive organs

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    • On the basis of symmetry:
      • Actinomorhic: A flower which can be divided by any plane into tow equal or similar halves
      • Zygomorphic: A flower which cannot be divided by any vertical plane into tow equal or similar halves.

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      • On the basis of floral parts arrangement
        • Hypogynous: The condition in which the sepals, petals and androecium are attached to the floral tube below the ovary.
        • Epigynous: The condition in which the sepals,petals and androecium are attached to the floral tube above the ovary
        • Perigynous: the condition in which the sepals, petals and andorecium are attached to the floral tube around the ovary.

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  • Calyx and corolla
    • No, aestivation, free or fused(poly or gamo), color, shape etc
    • Eg
      • Polysepalous: when sepals are free
      • Gamosepalous: when sepals are fused
  • Androecium
    • No, free or fused, extrose or introse
  • Gynoecium
    • Superior or inferior ovary, shape size of style and stigma, no of ovules

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Fruit

  • A fruit is regarded as mature or ripened ovary.
  • A fruit consist of two portions- pericarp (developed from the ovary) and seeds (developed from the ovules).
  • The pericarp consist of three parts
    • Epicarp (outer part)
    • Mesocarp( middle part)
    • Endocarp (inner part)

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  • Types of fruits
    • Berry: a fleshy fruit with succulent pericarp and many seeds embedding in the pulp. Eg Tomato
    • Capsule: Many seeded, one to many chambered dry furit. Eg Stramonium
    • Cremocarp: Two seeded fruit splitting longitudinally into two mericarps attached by carpophore eg Fennel
    • Drupe: A fleshy fruit with skiny epicarp, fleshy mesocarp and stony endocarp eg Mango
    • Hespiridium: A fleshy thick skinned berry with hard and leathery pericarp eg Citrus
    • Nut: A dry, indehicent, one seeded fruit with hard pericarp. Eg Cashew nut (kaaju)

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  • Shape:
    • Straight: No curving
    • Subcylindrical: Nearly cylindrical
    • Globular: globe shaped
    • Subshperical: nearly spherical
    • Ovoid: more or less oval shaped

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  • Adhesion
    • In superior fruit, the floral part or its remains is attached to the base eg Capsicum
    • In case of inferior fruit, the floral part is surmounted at its apex
  • Pericarp and its differentiation
    • Color, texture, markings (wrinkles, ridges)
  • Seeds
    • Number, shape, size, color, hilium, micropyle, testa etc

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Seeds

  • The fertilized ovule is known as seed.
  • Most of the seeds have two seed coats outer testa and inner tegmen.
  • The seed is attached to placenta by stalk or funicle.
  • The raphae is the ridge of fibrovascular tissues formed by the adhesion of funicle and testa.
  • Hilium is the scar left on the seed where it separates from the funicle.
  • Micropyle is the minute opening in the seed coat for the absorption of water.

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  • Shape size and color
    • Seed coats : thickness and texture
    • Hilium and micropyle: size and position
    • Perisperm and endosperm: presence or absence along with the nature of reserved food material.
    • Note: Perisperm is the nutritive material found outside the embroyo sac, is the remains of nucellus. Endosperm is the tissue that surrounds and nourishes the embroyo, developed from primary endosperm nucleus.

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Histology of different plant parts

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Root

  1. Epiblema
    • Epidermis of the root, usually single layered

2. Cortex

    • Consists of parenchymatous cells.
    • Innermost layer of cortex is called endodermis.

4. Stele

    • Consists of xylem, phloem, pericycle and pith.

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Stem

1. Epidermis

    • Single layer consisting of compactly arranged parenchymatous cells.

2. Cortex

a. Hypodermis

      • Collenchymatous, continuous ring or patches

b. General cortex

      • Parenchymatous , may contain starch grain, mucilage etc.

c. Endodermis

      • Innermost layer of cortex.

3. Stele

a. Pericycle

      • Composed of sclerenchymatous fibres, usually absent in monocot.

b. Vascular bundles

      • In dicot, it is arranged in ring form
      • In monocot, it is scattered.

c. Medullary rays

      • Radiating bands of parenchymatous cells which separate vascular bundles

d. Pith

      • Central part of stele.
      • May contain starch grains, mucilage, calcium oxalate crysals etc.

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Leaf

1. Epidermis

    • Usually single layered, composed of compactly arranged barrel shaped cells, cuticle stomata trichome present

2. Mesophyll

    • Differentiated into palisade cells and spongy cells
    • Pallisade cells are columnar and more or less compactly arranged at right angles to epidermis.
    • Spongy cells are oval shaped and contains abundant intercellular spaces.

3. Vascular bundles

    • Xylem is found towards upper epidermis while the band of phloem is found towards lower epidermis.
    • The bundle sheath is usually composed of parenchymatous cells.

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Secondary growth in root

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Root after secondary growth

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Secondary growth in stem

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Stem after secondary growth

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Stem after secondary growth

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Reference

  • http://www.pharmacy180.com/article/study-of-different-tissue-systems-19/