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How to ID Anemones in

Texas, OK, and southern NM

Compiled by R. S. Pfau�https://www.inaturalist.org/journal/pfau_tarleton/20891

Be sure to get pics of at least three features: (1) the stem-like stalk (including bracts) showing the region above and below the bract (to see if stalk below bract is pubescent or smooth), (2) the basal leaves (to see the pattern of division), (3) the flower. In western half of TX, also note if bracts are two-tiered, with two or more flowers growing from a single scape.

some leaves deeply divided multiple times

Suzette Rogers

scape smooth below bract (pubescent above)

Anemone caroliniana

Suzette Rogers

basal leaves

bracts�(also called involucres)

Jennifer Stockemer

scape

Anemone caroliniana

sepals (tepals; anemones lack petals)

scape pubescent below bract

Anemone berlandieri

rhizomes

Anemone berlandieri: All basal leaves lobed, dissected differently from bracts [leaves not typically deeply divided, lobed instead–unless infected with fungus]; bracts borne above middle of scape [ stem-like stalk] at anthesis [the period during which a flower is fully open and functional; this character is not accurate before/after anthesis]. Scape pubescent below bracts (varies as scape lengthens – longer scapes appear less fuzzy). Styles very short. Sepal color varies: white, pink, purple. No rhizomes. Despite common name, sepal number can exceed 10. Found across most of TX.

Anemone caroliniana: One or more basal leaves finely divided similarly to bracts [but early winter leaves are lobed like A. berlandieri]; bracts borne below middle of scape (often near ground) at anthesis [caution: the scape of other species elongates rapidly during anthesis and they can start to look like A. caroliniana in this regard]. Scape smooth below bracts [and pubescent above]. Styles very long. Sepals appear doubled (often twice the number of A. berlandieri–but this varies in A. berlandieri). Sepal color from white to deep purple. Rhizomes present. Found primarily in eastern half of TX, but reported from eastern TX panhandle (although these western populations may be an undescribed species–research on this is in progress by R. S. Pfau). Occurs only in sandy soils.

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Anemone okennonii: Known also as A. tuberosa var. texana. Often with two-tiered bracts. Leaves 2-3 ternate [three leaflets each divided 2-3 more times]. Basal and bract leaves dissimilar--bract leaves narrow (see slide 4). 7-11 sepals 3-4.5 mm wide. Sepal color white. Often with two-tiered bracts, with two or more flowers growing from a single scape. Distribution not well documented (appears to range from Val Verde Co. near Rio Grande north to perhaps Garza Co. and southeastern NM). http://plants.jstor.org/stable/10.5555/al.ap.flora.fna003_anemone_okennonii http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=101733

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Anemone edwardsiana

Alison Northup

joshua_tx

three broad leaflets, not greatly divided

all 3 species on this page often have two or more flowers growing from single scape (bracts two-tiered)

Anemone tuberosa: Bract leaves broad and highly divided–very similar to basal leaves (see slide 4). Often with two-tiered bracts. Styles pinkish in color. Sepal color varies from white to pink. In TX, found only near El Paso.

http://www.efloras.org/object_page.aspx?object_id=5538&flora_id=1

http://swbiodiversity.org/seinet/taxa/index.php?taxon=741

Anemone tuberosa

Descriptions modified largely from http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=101733, with additions from Shinners & Mahler's Flora of North Central Texas and personal observations.

Images provided (with permission) by iNaturalist observers Alison Northup, Suzette Rogers, Jennifer Stockemer, joshua_tx, and lonnyholmes.

Anemone edwardsiana: Often with two-tiered bracts, with two or more flowers growing from a single scape. Bract leaves wide. Basal leaves usually 1-ternate [three broad leaflets, not greatly divided] and 8-20 sepals 2-3 mm wide. Sepal color white. Can be difficult to distinguish from A. berlandieri unless two-tiered bracts are visible. Found only in limestone outcroppings of southern Edwards Plateau of TX.

Anemone okennonii

three leaflets, each deeply divided

bract leaves very narrow

bract leaves wide

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Anemone caroliniana

  • note long styles (arrow)

Anemone berlandieri

  • note short styles (arrow)

A. caroliniana vs. A. berlandieri

Floral details

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Anemone okennonii

  • Styles pinkish

Anemone edwardsiana

  • Styles, and overall fruit, green

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Anemone tuberosa vs. A. okennonii

Bract details

Anemone tuberosa

Anemone okennonii

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Undescribed species: Anemone 'pilosa' nom. prov.

-Pubescent below and above bracts, including lower surface of bracts and sepals (a)

-Bracts similar to A. berlandieri; single flower emerging from scape (a)

-"Doubled" sepals (b)

-Leaves highly divided (except early winter leaves) (c)

-Short styles (d)

-Sepals usually white, but purple variant exists

(a)

(b)

(c)

(d)

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Map from Ehrendorfer et al. (2009)

“Taxonomic revision, phylogenetics and transcontinental distribution of Anemone section Anemone (Ranunculaceae)”

This map (like most published maps, unfortunately) is a rough approximation. Especially regarding A. edwardsiana and A. okennonii. -RP

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Anemone phenology (records from Texas only, as of December 2022)

A. berlandieri

A. caroliniana

A. edwardsiana

A. okennonii

A. tuberosa