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1-4 Twitch

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Muscle Twitch in Frog Experiment

  • Threshold is minimum voltage necessary to produce action potential
    • a single brief stimulus at that voltage produces a quick cycle of contraction & relaxation called a twitch (lasting less than 1/10 second)
  • A single twitch contraction is not strong enough to do any useful work

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Twitch contraction

  • Phases of a twitch contraction
    • latent period (2 msec delay)
      • only internal tension is generated
      • no visible contraction occurs since �only elastic components are being �stretched
    • contraction phase
      • external tension develops as muscle �shortens
    • relaxation phase
      • loss of tension & return �to resting length as calcium returns to SR

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Recruitment & Stimulus Intensity

  • Stimulating the whole nerve with greater and greater voltage produces stronger contractions
  • More motor units are being recruited
    • called multiple motor unit summation
    • lift a glass of milk versus a whole gallon of milk

Maximal

recruitment

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Production of Variable Contraction Strengths�Twitch and Treppe Contractions

  • Stimulating a muscle at variable frequencies
    • low frequency (up to 10 stimuli/sec)
      • each stimulus produces an identical twitch response
    • moderate frequency (between 10-20 stimuli/sec)
      • each twitch has time to recover but develops more tension than the one before (treppe or staircase phenomenon)
        • calcium was not completely put back into SR
        • heat of tissue increases myosin ATPase effeciency (warm-up exercises)

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Production of Variable Contraction Strengths�Incomplete and Complete Tetanus

  • Higher frequency stimulation (20-40 stimuli/second) generates gradually more strength of contraction
    • each stimuli arrives before last one recovers
      • temporal summation or wave summation
    • incomplete tetanus = sustained fluttering contractions
  • Maximum frequency stimulation (40-50 stimuli/second)
    • muscle has no time to relax at all
    • twitches fuse into smooth, prolonged contraction called complete tetanus
    • rarely occurs in the body