1 of 59

Plant Form and Function

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2 of 59

Introduction: vegetative plant parts

Section 22.1

If asked to picture a plant, you probably wouldn’t think of unusual examples like a Venus flytrap or a barrel cactus.

Figure 22.3

Carnivory (Venus flytrap)

©Win Initiative/Getty Images RF

Water storage (cactus)

©G.C. Kelly/Science Source

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Vegetative plant parts

Section 22.1

But like other flowering plants, these have roots, stems, leaves, flowers, fruits, and seeds.

Figure 22.3

Carnivory (Venus flytrap)

©Win Initiative/Getty Images RF

Water storage (cactus)

©G.C. Kelly/Science Source

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Vegetative plant parts: modified leaves

Section 22.1

Harsh environments have selected for adaptions in these plants, such as modified leaves.

Figure 22.3

Modified leaves

Pollinator attraction (poinsettia)

©Design Pics/Don Hammond RF

Carnivory (Venus flytrap)

©Win Initiative/Getty Images RF

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Vegetative plant parts = nonreproductive plant parts

Section 22.1

This chapter explores the anatomy and physiology of vegetative (nonreproductive) plant parts.

Figure 22.3

Carnivory (Venus flytrap)

©Win Initiative/Getty Images RF

Water storage (cactus)

©G.C.Kelly/Science Source

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Naming the vegetative plant parts

Section 22.1

Vegetative plant parts include stems, leaves, and roots. These organs work together.

Figure 22.1

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Vegetative plant parts: the shoot

Section 22.1

The shoot is the aboveground part of the plant.

Figure 22.1

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Vegetative plant parts: the stem

Section 22.1

The shoot’s stem supports the leaves, which produce carbohydrates by photosynthesis.

Figure 22.1

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Vegetative plant parts: the roots

Section 22.1

Some of the sugar produced in the shoot system travels through the stem to the roots, which are usually below ground.

Figure 22.1

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Vegetative plant parts: functions of roots

Section 22.1

Roots anchor the plant and absorb water and minerals that move via the stem to the leaves.

Later, we will explore how water, minerals, and sugars travel through plants.

Figure 22.1

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Vegetative plant parts: nodes and internodes

Section 22.1

Leaves attach to stems at nodes. Spaces between nodes are internodes.

Figure 22.1

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Vegetative plant parts: axillary buds

Section 22.1

Each node also features an axillary bud, an undeveloped shoot that could form

a new branch or flower.

Figure 22.1

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Two types of plants

Section 22.1

Biologists divide plants into two categories based on the characteristics of the stem.

Figures 22.1, 24.4

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Herbaceous and woody plants

Section 22.1

A herbaceous plant has a green, soft stem.

A woody plant is made of tough, bark-covered wood.

Figures 22.1, 24.4

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Vegetative plant parts: specialized stems

Section 22.1

Natural selection produces stems, leaves, and roots with various forms.

Figure 22.3

  1. Specialized stems

Climbing (grape tendrils)

Underground nutrient storage (iris rhizomes)

Water storage (cactus)

Defense

(honey locust thorns)

(a, vine): ©Franz Krenn/Science Source; (a, iris): ©Dwight Kuhn; (a, cactus): ©G.C. Kelly/Science Source; (a, thorns): ©Kenneth W. Fink/Science Source

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Vegetative plant parts: specialized leaves

Section 22.1

Natural selection produces stems, leaves, and roots with various forms.

Figure 22.3

  1. Specialized leaves

Nutrient storage (onion)

Pollinator attraction (poinsettia)

Carnivory (Venus flytrap)

Asexual reproduction (kalanchöe)

(b, onion): ©YAY Media AS/Alamy RF; (b, poinsettia): ©Design Pics/Don Hammond RF; (b, flytrap): ©Win Initiative/Getty Images RF; (b, kalanchoe)

22-16

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Vegetative plant parts: specialized roots

Section 22.1

Natural selection produces stems, leaves, and roots with various forms.

Figure 22.3

  1. Specialized roots

Nutrient storage (carrot)

Oxygen absorption (mangrove trees)

Photosynthesis (orchid aerial roots)

Support

(prop roots of screw pine)

(c, carrots): ©Huw Jones/Photolibrary/Getty Images; (c, mangrove): ©Tim Laman/Getty Images RF; (c, orchid): ©Settawut Visedbubpha/123R; (c, screw pine): ©Steven P. Lynch/McGraw-Hill Education

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Plant cells build tissues

Section 22.3

We’ve seen the organs and organ systems of plants. Now let’s zoom in and learn about the cells and tissues that make up these organs.

Figure 22.1

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Three main tissue types

Section 22.2

Plants have three main tissue types:

  • Ground tissue makes up most of the plant body.
  • Vascular tissues (xylem and phloem) transport materials within the plant.
  • Dermal tissue covers the plant.

Figure 22.6

22-19

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Ground tissue

Section 22.2

Ground tissue consists of three main cell types: parenchyma, collenchyma, and sclerenchyma.

Figure 22.4

Cell Type

Description

Alive at Maturity

Functions

Parenchyma

  • Most abundant cell type in primary plant body
  • Thin primary cell walls
  • Unspecialized
  • Can divide at maturity

Yes

Make up most nonwoody tissues; carry out photosynthesis, respiration, gas exchange, secretion, wound repair, and storage

Collenchyma

  • Elongated cells
  • Unevenly thickened primary cell walls

Yes

Elastic support for growing stems and leaves

Sclerenchyma:

Fiber

  • Long, slender cells
  • Thick secondary cell walls high in lignin

No

Inelastic support for nongrowing plant parts

Sclerenchyma:

Sclereid

  • Variable shapes, generally not elongated
  • Thick secondary cell walls high in lignin

No

Inelastic support for nongrowing plant parts

(parenchyma): ©Malcolm Park microimages/Alamy; (collenchyma): ©Biophoto Associates/Science Source; (fibers): ©Steven P. Lynch/McGraw-Hill Education; (sclereid): ©Garry Delong/Oxford Scientific/Getty Images

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Functions of ground tissue

Section 22.2

The cells that compose ground tissue are important sites of photosynthesis, respiration, storage, and support.

Figure 22.4

Cell Type

Description

Alive at Maturity

Functions

Parenchyma

  • Most abundant cell type in primary plant body
  • Thin primary cell walls
  • Unspecialized
  • Can divide at maturity

Yes

Make up most nonwoody tissues; carry out photosynthesis, respiration, gas exchange, secretion, wound repair, and storage

Collenchyma

  • Elongated cells
  • Unevenly thickened primary cell walls

Yes

Elastic support for growing stems and leaves

Sclerenchyma:

Fiber

  • Long, slender cells
  • Thick secondary cell walls high in lignin

No

Inelastic support for nongrowing plant parts

Sclerenchyma:

Sclereid

  • Variable shapes, generally not elongated
  • Thick secondary cell walls high in lignin

No

Inelastic support for nongrowing plant parts

(parenchyma): ©Malcolm Park microimages/Alamy; (collenchyma): ©Biophoto Associates/Science Source; (fibers): ©Steven P. Lynch/McGraw-Hill Education; (sclereid): ©Garry Delong/Oxford Scientific/Getty Images

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22 of 59

Vascular tissue

Section 22.2

Vascular tissues transport water, minerals, carbohydrates, and other dissolved compounds.

Figure 22.5

Cell Type

Functions

XYLEM

Tracheid

Conduct water and minerals through pits

Vessel element

Conduct water and minerals through pits and perforated end walls

PHLOEM

Sieve tube element

Conduct dissolved sucrose and other organic compounds through sieve plates

Companion cell

Transfer materials into and out of

sieve tube elements

©Biophoto Associates/Science Source

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Vascular tissue: xylem

Section 22.2

Xylem tissue transports water and minerals from the roots to other plant parts. It consists of long, narrow cells called tracheids and wide, barrel-shaped cells called vessel elements.

Figure 22.5

Cell Type

Functions

XYLEM

Tracheid

Conduct water and minerals through pits

Vessel element

Conduct water and minerals through pits and perforated end walls

PHLOEM

Sieve tube element

Conduct dissolved sucrose and other organic compounds through sieve plates

Companion cell

Transfer materials into and out of

sieve tube elements

©Biophoto Associates/Science Source

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Vascular tissue: phloem

Section 22.2

Phloem tissue transports dissolved organic compounds like sugars. Sieve tube elements are the conducting cells; they are separated by sieve plates.

Figure 22.5

Cell Type

Functions

XYLEM

Tracheid

Conduct water and minerals through pits

Vessel element

Conduct water and minerals through pits and perforated end walls

PHLOEM

Sieve tube element

Conduct dissolved sucrose and other organic compounds through sieve plates

Companion cell

Transfer materials into and out of

sieve tube elements

©Biophoto Associates/Science Source

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Vascular tissue: phloem cells

Section 22.2

Companion cells transfer materials in and out of sieve tubes.

Figure 22.5

Cell Type

Functions

XYLEM

Tracheid

Conduct water and minerals through pits

Vessel element

Conduct water and minerals through pits and perforated end walls

PHLOEM

Sieve tube element

Conduct dissolved sucrose and other organic compounds through sieve plates

Companion cell

Transfer materials into and out of

sieve tube elements

©Biophoto Associates/Science Source

22-25

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Dermal tissues

Section 22.2

Dermal tissue covers the plant; it consists of the epidermis, which is coated with a waxy cuticle.

Figure 22.7

©Steven P. Lynch/McGraw-Hill Education

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Cuticle and stomata

Section 22.2

The cuticle conserves water and protects the plant. Pores in the cuticle, called stomata, allow leaves to exchange gases with the atmosphere.

Figure 22.7

©Steven P. Lynch/McGraw-Hill Education

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Guard cells

Section 22.2

Guard cells surround each stoma and control its opening and closing.

Figure 22.8

©Steven P. Lynch/McGraw-Hill Education

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29 of 59

Tissues build stems, leaves, and roots

Section 22.3

The three tissue types make up the stems, leaves, and roots of the plant.

Let’s look at each of these organs, starting with the stem.

Figure 22.6

22-29

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Tissue types found in a stem

Section 22.3

Ground tissue occupies most of the stem of a herbaceous plant.

Vascular bundles are embedded in the ground tissue.

Dermal tissue covers the stem.

Figure 22.6

22-30

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31 of 59

Arrangement of tissues differs in monocots and eudicots

Section 22.3

Monocots and eudicots have different arrangements of vascular tissue and ground tissue in their stems.

Figure 22.9

  1. Corn (monocot)
  1. Sunflower (eudicot)

(a, stem): ©Steven P. Lynch/RF; (a, corn close up; b, both): ©Steven P. Lynch/McGraw-Hill Education

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Vascular bundles in monocot and eudicot stems

Section 22.3

In monocots, vascular bundles are scattered throughout the stem.

In eudicots, vascular bundles are arranged in a ring near the epidermis.

The cortex is ground tissue that fills the space between the epidermis and vascular bundles. The pith occupies the center of the stem.

Figure 22.9

  1. Corn (monocot)
  1. Sunflower (eudicot)

(a, stem): ©Steven P. Lynch/RF; (a, corn close up; b, both): ©Steven P. Lynch/McGraw-Hill Education

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33 of 59

Tissues found in leaves

Section 22.3

Ground tissue occupies most of a leaf.

Vascular bundles are embedded in the ground tissue.

Dermal tissue covers the leaf.

Figure 22.6

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The structure of leaves

Section 22.3

Leaves are flattened blades supported with a stalklike petiole.

Figure 22.10

Simple leaf

Compound leaves

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Simple vs. compound leaves

Section 22.3

Simple leaves have undivided blades.

Compound leaves are divided into leaflets attached to one petiole.

Figure 22.10

Simple leaf

Compound leaves

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Vein patterns on monocot and eudicot leaves

Section 22.3

Veins are vascular bundles inside leaves. Many monocots have parallel veins; most eudicots have netted veins.

Figure 22.11

Veins on typical monocot leaf

Veins on typical eudicot leaf

(both): ©Dwight Kuhn

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Mesophyll: the middle of a leaf

Section 22.3

Leaf anatomy shown here is that of a eudicot plant.

The ground tissue inside a leaf is called mesophyll, which consists of cells with abundant chloroplasts that produce sugars by photosynthesis.

Figure 22.12

©M. I. Walker/Science Source

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Stomata: locations of gas exchange

Section 22.3

When stomata are open, mesophyll cells exchange gases with the atmosphere.

Figure 22.12

©M. I. Walker/Science Source

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Mesophyll cells interact with vascular tissue

Section 22.3

Mesophyll cells also exchange materials with vascular tissues.

Figure 22.12

©M. I. Walker/Science Source

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40 of 59

Monocot leaf structures

Section 22.3

Monocots have similar leaf anatomy to dicots. Note the prominent bundle sheath cells in this monocot leaf, surrounded by a layer of mesophyll.

Figure 22.12

©Steven P. Lynch/McGraw-Hill Education

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Tissues of the root

Section 22.3

In a root, ground tissue surrounds a central core of vascular tissue.

Dermal tissue forms the root epidermis.

Figure 22.6

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Fibrous roots vs. taproots

Section 22.3

Roots might form a fibrous root system or a taproot system.

  • Fibrous roots are slender, shallow, and arise from the base of the stem.
  • Taproots are thick, deep, and have fewer branches than fibrous roots.

Figure 22.13

  1. Barley (monocot)
  1. Buttercup (eudicot)

(a): ©AI Telser/McGraw-Hill Education; (b): ©Ed Reschke/Photolibrary/Getty Images

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Root hairs

Section 22.3

Near each root’s tip, root hairs are extensions of the epidermis that

absorb water and minerals.

Figure 22.14

©Dr. Jeremy Burgess/Science Source

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44 of 59

Plants have flexible growth patterns

Section 22.4

Some plants never stop growing. These plants have indeterminate growth.

Plants that stop growing when they reach their mature size have determinate growth.

Table 22.3

TABLE 22.3 Meristem Types: A Summary

Type

Locations

Function

Apical

Terminal and axillary buds of shoots; root tips

Produces tissues that lengthen the tips of shoots and roots

Lateral

Internal cylinder along the length of roots and stems of woody plants

Thickens roots and stems

Intercalary

Between nodes of mature stems in grasses and other monocots

Regrowth of tissue if tip of stem is removed

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Meristems

Section 22.4

Plants grow by adding units, or modules, consisting of repeated nodes and internodes. Growth occurs at meristems, regions of active cell division.

Table 22.3

22-45

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Apical meristems

Section 22.4

Apical meristems produce tissues that lengthen the tips of shoots and roots.

Table 22.3

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Primary growth

Section 22.4

Primary growth occurs at the apical meristems. New cells can differentiate into any tissue type.

Figs. 22.15, 22.16

©Steven P.Lynch/McGraw-Hill Education

Apical meristems

©Steven P.Lynch/McGraw-Hill Education

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48 of 59

Intercalary meristems

Section 22.4

Intercalary meristems occur at the base of a leaf blade. �Grasses tolerate grazing because they have intercalary meristems that regrow a leaf from its base when the tip is munched off.

Table 22.3

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Secondary growth

Section 22.4

Secondary growth thickens roots and stems; this growth occurs at lateral meristems.

Table 22.3

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50 of 59

Secondary growth in woody plants

Section 22.4

Secondary growth occurs in woody plants. Two types of lateral meristems produce wood and bark:

  • Vascular cambium
  • Cork cambium

Figure 22.17

  1. Primary growth
  1. Secondary growth

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Vascular cambium

Section 22.4

The vascular cambium (highlighted green) produces secondary xylem toward the inside of the stem and secondary phloem toward the outside.

Figure 22.17

  1. Primary growth
  1. Secondary growth

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Wood

Section 22.4

Secondary xylem is more commonly called wood.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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Rays

Section 22.4

The vascular cambium also produces rays (highlighted with yellow), bands of parenchyma that extend from the center of the stem or root and transport nutrients laterally.

Figure 22.17

  1. Primary growth
  1. Secondary growth

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Bark

Section 22.4

Secondary growth produces bark, a collective term for all tissues outside of the vascular cambium.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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Cork cambium

Section 22.4

The cork cambium (highlighted white) produces parenchyma cells toward the inside and dense, waxy cells called cork toward the outside.

Cork is the outer protective layer of bark.

Figure 22.17

  1. Primary growth
  1. Secondary growth

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Periderm

Section 22.4

Together, the cork cambium, parenchyma cells, and nonliving cork make up the periderm, a protective layer that covers a woody stem or root.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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Heartwood

Section 22.4

Secondary xylem eventually becomes unable to conduct water, forming heartwood.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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Sapwood

Section 22.4

The lighter sapwood transports water and dissolved minerals.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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Tree rings

Section 22.4

Tree rings arise from alternating moist and dry seasons. Wood that forms in the spring has larger cells than wood that forms in the summer.

Figure 22.18

(b): ©Siede Preis/Getty Images RF; (c): ©Herve Conge/Phototake

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