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bioinformatics.ca

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Introduction to Spatial�Single Cell RNA-seq Technologies

Trevor Pugh, PhD, FACMG

Introductory Spatial 'Omics Analysis

Feb 20-21, 2025

bioinformatics.ca

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Learning Objectives

  1. Understand the conceptual shift in moving from bulk to single cell profiling
  2. Learn types, parameters and trade-offs of various single cell technologies
  3. Using cancer as an example, be exposed to scientific questions and experimental designs utilizing single cell analysis
  4. Become acquainted with advances and applications of spatial transcriptomics in brain research

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We are all made of cells: tissues consist of immune, stromal & many other cell types that interact physically and functionally

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Single cell analysis is not new…the revolution is in the scale, completeness, & quantitative nature of genomic technologies

One karyotype in one cell

In situ hybridization of one transcript

Quantification of all chromsomes in all cells

Visualization of 1,000s of genes expressed in all cells

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Single-cell analysis reveals heterogeneity in molecular profiles�at resolution bulk analysis may not permit

e.g. Six cells with heterogeneous

expression of three genes

Bulk analysis detects uniform

expression of all three genes

Single-cell analysis directly

measures diversity of expression

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“A wide variety of single-cell methods have now been developed to measure a broad range of cellular parameters”

Stuart and Satija. Nature Reviews Genetics. 20:257-272. May 2019

We will focus here

Lineage

State

Trajectory

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“Exponential scaling of single-cell RNA-seq in the past decade”

Svensson, Vento-Tormo, and Teichmann. Nat Protoc. 2018 Apr;13(4):599-604.

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New spatial technologies enable additional cellular metadata describing physical distances between cell types and cell states

https://www.10xgenomics.com/in-situ-technology

Stuart and Satija. Nature Reviews Genetics. 20:257-272. May 2019

10X Genomics

Xenium

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Cell organization & cell state is now measurable at all biological scales:�cells, tissues, organisms, populations

Du, Yang et al. J Transl Med. 2023; 21: 330.

https://www.10xgenomics.com/products/xenium-panels

Mouse Tissue Atlassing (379 genes)

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Several strategies to localize a transcript within a cell�a cell within a tissue; and a tissue within an organism

Rao, A., Barkley, D., França, G.S. et al. Exploring tissue architecture using spatial transcriptomics. Nature 596, 211–220 (2021).

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10X Genomics as an example of commercialized spatial platforms that include droplets, sequencing, and probe panels

https://omicsomics.blogspot.com/2021/02/more-details-on-10xs-sample-profiling.html

Janesick et al. bioRxiv 2022.10.06.510405 doi: https://doi.org/10.1101/2022.10.06.510405

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How do you choose a spatial transcriptomic method?

Gene throughput

Unbiased sequence everything versus targeted panels?

Sequence information

Do you need knowledge of mutations, fusions, splice variants?

Sensitivity

Targeted approaches more sensitive than sequencing-based methods

Resolution

Limitations of spotted arrays versus optical diffraction limit (subcellular)

Area size

Small 6-10mm2 areas versus full-slide imaging. Trade-off with time.

Feasibility

Access to tissues, imaging, panels, bioinformatics and reference data

Summarized from Rao, A., Barkley, D., França, G.S. et al. Nature 596, 211–220 (2021).

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Cancer as an example

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An example: glioblastomas contain self-renewing cancer stem cells that underlie tumour initiation & therapeutic resistance

Richards, Whitley et al. Nature Cancer. 2021 Feb. Gradient of developmental and injury-response transcriptional states defines functional vulnerabilities underpinning glioblastoma heterogeneity

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Brain tumour stem cell cultures derived from primary GBMs

Single cell RNA-seq of

all cells within a tumour

Single cell RNA-seq of

only the brain tumour stem cells

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scVelo. Bergen et al.

Nat Biotechnol 38, 1408–1414 (2020).

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2,000 um

Glioblastoma

Grade IV

Primary – Surgical specimen

Formalin-fixed paraffin embedded tissue

Applied custom 300 gene probe panel to profile transcriptional cancer stem cell gradients in GBM

(300 gene immune panel also available)

x position (um)

y position (um)

Shamini Ayyadhury,

Gary Bader, Peter Dirks

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209,944 cells detected

266 transcripts

per cell

Transcripts per bin

Size (um2)

Cell Size Distribution

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1,000

4k

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Genes

per cell

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Genes

95.9% transcripts within cells

0.308 cells per 100 um2

93 genes from

266 transcripts

per cell

x position (um)

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Transcriptional cluster

2,000 um

x position (um)

Likely blood vessels

(COL1A1, COL3A1, COL4A1, PDGFRB, ACTA2, TGBFI)

Likely immune cells

(CXCL8, HLAs, RGS2, ITGAX, TLR2)

Shamini Ayyadhury,

Gary Bader, Peter Dirks

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Highly specific cell populations within all types of cancer characterized over the past 3 years during rapid adoption of spatial transcriptomics

Cilento, M.A., Sweeney, C.J. & Butler, L.M J Cancer Res Clin Oncol 150, 296 (2024)

Spatial transcriptomics in cancer research and potential clinical impact: a narrative review.

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Single cell RNA-seq reference sets from all parts of the brain, healthy and diseased

Piwecka, M., Rajewsky, N. & Rybak-Wolf, A. Single-cell and spatial transcriptomics: deciphering brain complexity in health and disease. Nat Rev Neurol 19, 346–362 (2023).

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The Allen Brain Map is a powerful source of reference data, biological validation resources, & computational tools for brain research

https://portal.brain-map.org

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Revisiting the Learning Objectives

  1. A plethora of single cell technologies have opened windows into cell biology that were closed to bulk approaches that “average out” signal
  2. The same biology may be measurable using multiple methods 🡪 tailor experimental approaches to specific scientific questions answerable by available samples & technologies
  3. Multiple cellular components can be queried from one single cell experiment, e.g. immune & cancer cells inhabiting tumours
  4. Consider spatial technologies and compatible reference sets based on gene throughput, sequence information, sensitivity, resolution, area size, and feasibility required to pursue your scientific goals