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What plausible hypothesis could account for the origin of life?

What intermediate stages could there have been between non-living matter and the first living cells?

What are the 3 principles of cell theory?

A2.1 Origin of Cells

Guiding Questions

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A2.1 Origin of Cells

Can science explain the origin of life?

Guiding question

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A2.1.1—Conditions on early Earth and the pre-biotic formation of carbon compounds

Include the lack of free oxygen and therefore ozone, higher concentrations of carbon dioxide and methane, resulting in higher temperatures and ultraviolet light penetration. The conditions may have caused a variety of carbon compounds to form spontaneously by chemical processes that do not now occur

A2.1.2—Cells as the smallest units of self-sustaining life

Discuss the differences between something that is living and something that is non-living. Include reasons that viruses are considered to be non-living.

A2.1.3—Challenge of explaining the spontaneous origin of cells

Cells are highly complex structures that can currently only be produced by division of pre-existing cells. Students should be aware that catalysis, self-replication of molecules, self-assembly and the emergence of compartmentalization were necessary requirements for the evolution of the first cells.

A2.1.4—Evidence for the origin of carbon compounds

Evaluate the Miller–Urey experiment.

A2.1.5—Spontaneous formation of vesicles by coalescence of fatty acids into spherical bilayers

Formation of a membrane-bound compartment is needed to allow internal chemistry to become different from that outside the compartment

A2.1.6—RNA as a presumed first genetic material

RNA can be replicated and has some catalytic activity so it may have acted initially as both the genetic material and the enzymes of the earliest cells. Ribozymes in the ribosome are still used to catalyse peptide bond formation during protein synthesis

A2.1.7—Evidence for a last universal common ancestor

Students should develop an appreciation of the immense length of time over which life has been evolving on Earth

A2.1.8—Approaches used to estimate dates of the first living cells and the last universal common ancestor

Students should develop an appreciation of the immense length of time over which life has been evolving on Earth

A2.1.9—Evidence for the evolution of the last universal common ancestor in the vicinity of hydrothermal vents

Include fossilized evidence of life from ancient seafloor hydrothermal vent precipitates and evidence of conserved sequences from genomic analysis.

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So… what came first…?

A2.1 Origin of Cells

Justify your answer

A2.1.4 Evidence for the origin of carbon compounds

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List things we would need to create life on Mars

A2.1 Origin of Cells

Can we live on Mars? Justify your answer

A2.1.4 Evidence for the origin of carbon compounds

Challenge: What processes and/or molecules would need to be present? How do we create life?

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A2.1 Origin of Cells

What are the principles of cell theory?

A2.1.2—Cells as the smallest units of self-sustaining life

1.  All living things are composed of cells (or cell products)

2.  Cells are the smallest unit of life

3.  Cells only arise from pre-existing cells

Challenge: What are the exceptions to the cell theory and why?

Challenge: Why are viruses consdered to be non-living?

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All cells have:

  1. membrane compartment
  2. genetic material
  3. metabolic processes

A2.1 Origin of Cells

What do we know about cells?

A2.1.1—Conditions on early Earth and the pre-biotic formation of carbon compounds

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A2.1 Origin of Cells

What are the principles of cell theory?

A2.1.2—Cells as the smallest units of self-sustaining life

1. Cells are highly complex structures and no mechanism has been found for producing cells from simpler subunits.

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What are the principles of cell theory?

2. All known examples of growth be it of a tissue, an organism or a population, are all a result of cell division.

A2.1 Origin of Cells

A2.1.2—Cells as the smallest units of self-sustaining life

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What are the principles of cell theory?

3. Viruses are produced from simpler subunits, but they do not consist of cells, and they can only be produced inside the host cells that they have infected.

A2.1 Origin of Cells

A2.1.2—Cells as the smallest units of self-sustaining life

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What are the principles of cell theory?

4. Genetic code is universal each of the 64 codons (a codon is a combination of 3 DNA bases) produces the same amino acid in translation, regardless of the organism

The logical deduction is that all cells have arisen as the result of cell division from a single common ancestor.

A2.1 Origin of Cells

A2.1.6—RNA as a presumed first genetic material

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A2.1 Origin of Cells

Can we live on Mars? Justify your answer

A2.1.4 Evidence for the origin of carbon compounds

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A2.1 Origin of Cells

What was early Earth like 4.7 billion years ago?

A2.1.4 Evidence for the origin of carbon compounds

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A2.1 Origin of Cells

What was early Earth like 4.7 billion years ago?

A2.1.4 Evidence for the origin of carbon compounds

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Ozone (O3) is a form of oxygen that is found in the stratosphere, a region of the atmosphere located approximately 10 to 50 kilometers.

The ozone is controlled by the presence (or lack) of oxygen molecules.

A2.1 Origin of Cells

Suggest reasons why the presence of oxygen has affected life on Earth.

A2.1.4 Evidence for the origin of carbon compounds

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  • UV causes mutations in genetic material (DNA/RNA).
  • Ozone limited the amount of UV entering
  • CO2 trapped the heat in, to warm the Earth

A2.1 Origin of Cells

Suggest reasons why the presence of oxygen has affected life on Earth.

A2.1.4 Evidence for the origin of carbon compounds

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1. The non-living synthesis of simple organic molecules (from inorganic compounds) such as sugars and amino acids

2. The assembly of these organic molecules into polymers

3. The formation of plasma membranes

4. DNA and enzymes (RNA can replicate, catalyse reactions and act as instructions)

A2.1 Origin of Cells

Four processes needed for spontaneous life on Earth

A2.1.1—Conditions on early Earth and the pre-biotic formation of carbon compounds

Chemical Evolution

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A2.1 Origin of Cells

From these 4 requirements for life.. which came first? why?

A2.1.3—Challenge of explaining the spontaneous origin of cells

Plasma membranes (Protocells)

Genes RNA can replicate

Metabolism catalyse reactions

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Organic synthesis requires the presence of a range of inorganic molecules and an energy source to combine them into organic forms

A2.1 Origin of Cells

Where did the inorganic molecules and energy come from?

A2.1.4 Evidence for the origin of carbon compounds

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  • Comets and meteorites contain a variety of organic molecules (~20% of a comet's tail is comprised of organic material)
  • Heavy bombardment about 4,000 million years ago may have delivered both organic compounds and water to early Earth

A2.1 Origin of Cells

A2.1.4 Evidence for the origin of carbon compounds

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Hydrothermal vents: Regions of the ocean floor where superheated water is released from the Earth's crust

A2.1 Origin of Cells

A2.1.4 Evidence for the origin of carbon compounds

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Volcanoes: Intensive volcanic activity could provide sufficient thermal activity either on land or under the seabed

A2.1 Origin of Cells

A2.1.4 Evidence for the origin of carbon compounds

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Extraterrestrial locations: Other planets (e.g. Mars) may have been subjected to appropriate conditions (compound transfer via meteorites)

A2.1 Origin of Cells

A2.1.4 Evidence for the origin of carbon compounds

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Miller and Urey recreated the conditions of pre-biotic Earth in a closed system consisting of a series of flasks and tubes

A2.1 Origin of Cells

What did Miller and Urey do?

A2.1.4—Evidence for the origin of carbon compounds

Methane & Ammonia

Water (Ocean)

Organic Molecules were synthesised

Amino Acids, Lipids and Sugars

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Miller and Urey’s experiment:�- used high levels of methane (CH4) which was actually in relatively low abundance

-used Electrical discharges instead of UV light

A2.1 Origin of Cells

What did Miller and Urey do?

A2.1.4—Evidence for the origin of carbon compounds

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Weak Hydrophobic Interactions enables the cell membrane to be fluid and flexible.

B2.1.13—Membrane fluidity and the fusion and formation of vesicles

B2.1 Membranes and Membrane Transport

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Breaking and building the cell membrane requires ATP.

B2.1.13—Membrane fluidity and the fusion and formation of vesicles

B2.1 Membranes and Membrane Transport

Endocytosis

Exocytosis

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Pinocytosis - Microvilli

Phagocytosis - Immune Response

RM Endocytosis - Cholesterol for Membrane Synthesis

B2.1.13—Membrane fluidity and the fusion and formation of vesicles

B2.1 Membranes and Membrane Transport

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B2.2.9—Structure and function of vesicles in cells

B2.2 Organelles and compartmentalization

Clathrin

Membrane curvature

Clathrin Lattice Formed

Clathrin Coated Vesicle

Clathrin Uncoating

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Transport vesicles, lysosomes and peroxisomes usually contain a single membrane. Phagolysosomes are usually double-membrane.

B2.2.9—Structure and function of vesicles in cells

B2.2 Organelles and compartmentalization

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4. Formation of membranes to package the organic molecules

Experiments have shown that phospholipids natural assemble into bilayers, if conditions are correct.

Formation of the bilayer creates an isolated internal environment.

The formation of an internal environment means that optimal conditions, e.g. for replication or catalysis can be maintained.

A2.1 Origin of Cells

Four processes needed for spontaneous life on Earth

A2.1.5—Spontaneous formation of vesicles by coalescence of fatty acids into spherical bilayers

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Coalescence is the process where two or more oil droplets merge together to form a larger oil droplet.

A2.1 Origin of Cells

How do vesicles interact?

A2.1.5—Spontaneous formation of vesicles by coalescence of fatty acids into spherical bilayers

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A2.1 Origin of Cells

What is the central dogma?

A2.1.6—RNA as a presumed first genetic material

Ribozymes -

act as catalysts

Proteins (enzymes)

contains genetic material and catalyst to make more proteins

RNA

RNA molecules that have catalytic properties

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  • Universal Genetic Code (central dogma)
  • Metabolic Pathways (glycolysis)
  • Molecular Structure (ribosomes)
  • Phylogeny (DNA and Amino Acid sequencing)
  • Fossils (clay near hydrothermal vents)

A2.1 Origin of Cells

A2.1.7—Evidence for a last universal common ancestor (LUCA)

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  • Geological Evidence - studying the minerals present in certain rocks
  • Genomic Clocks - comparing differences and estimating the amount of time between required for mutations and variation to occur

A2.1 Origin of Cells

A2.1.8—Approaches used to estimate dates of the first living cells and the last universal common ancestor

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A2.1 Origin of Cells

A2.1.9—Evidence for the evolution of the last universal common ancestor in the vicinity of hydrothermal vents

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  • Nuvvuagittuq greenstone belt
  • Slithers of rocks contain bacteria dated from 3.77-4.28 billion years
  • Similar structures to modern day prokaryotes
  • Iron-oxidising (removes electrons for metabolism)

A2.1 Origin of Cells

A2.1.9—Evidence for the evolution of the last universal common ancestor in the vicinity of hydrothermal vents

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Watch to 1 min 40 secs:

A2.1 Origin of Cells

Spontaneous Generation

1.5.U2 The first cells must have arisen from non-living material.

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But when Pasteur broke the swan neck off the bottle microorganisms soon grew in the broth.

A2.1 Origin of Cells

Pasteur’s Swan Neck Flask Experiment

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Estimating the dates of the first living cells and the Last Universal Common Ancestor (LUCA) is a challenging task due to the limited availability of direct evidence from the distant past.

  • Fossil Record
  • Molecular Clocks
  • Comparative Genomics
  • Phylogenetic Analysis
  • Geological and Geochemical Evidence

A2.1 Origin of Cells

What did Miller and Urey do?

A2.1.8 Approaches used to estimate dates of the first living cells and the last universal common ancestor