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CHS 9 - MUSCLE & MUSCLE TISSUE

Functions of Muscles:

Movement - pulling on something to change its position

Maintaining Posture - tension to prevent movement, keeping vertebral column upright

Stabilizing Joints - attaching to bone & keeping them close together

Heat Generation - release of energy during metabolism

Functional Characteristics of Muscles:

Extensibility - ability to be stretched or extended

Excitability/Irritability - ability to receive and respond to stimuli

Contractility - ability to shorten forcibly when adequately stimulated

Elasticity - ability to resume its resting length after being stretched

Types of Muscle Tissue:

Cardiac Muscles -

Cells have branching chains of uni or binucleate striations, intercalated

discs

Involuntary with intrinsic system regulation, hormones, and autonomic

nervous system controls

Found in the heart

Skeletal Muscles - (most of Ch. 9 is about)

Single, very long, multinucleated cells with striations

Voluntary via axonal endings of the somatic nervous system

Attached to bones and skin

Powerful

Smooth Muscles -

Single, fusiform, uninucleate, no striations

Involuntary with autonomic nerves, hormones, and local chemicals

Found on the wall of hollow organs (stomach, bladder, airways) and tracts

Fibers are small and spindle shaped

Types of Muscle Contractions:

Isometric - tension continues to increase but the muscle neither shortens nor

lengthens

Isotonic - the muscle changes in length and tension remains constant through most

of the contractile period

Concentric - the muscle shortens and does work

Eccentric - the muscle contracts as it lengthens

Types of Muscle Chemicals:

Lactic Acid - product of anaerobic glycolysis, increases during repetitive muscle

contractions, and causes muscle fatigue

Myoglobin - oxygen storage molecules in muscle

Creatine Phosphate - high-energy compound in muscle

Acetylcholine - neurotransmitter at the neuromuscular junction

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Acetylcholinesterase - enzyme released into neuromuscular junction to break down

acetylcholine

Calmodulin - cytoplasmic, calcium-binding protein

Ca++ - an ion released by terminal cisternae into the sarcoplasm to bind with troponin to

remove the blocking action of tropomyosin, resulting in cross bridges

Triad - structure in a skeletal muscle fiber made up of TWO terminal cisterna and a

transverse tubule

Elastic Filaments - allows the muscle fiber to recoil when the contraction ends

Types of Proteins and Their Actions:

Myosin - changes shape during the contraction cycle

In sliding filament model, myofilaments slide over one another resulting in the

overlapping of myosin with actin

Tropomyosin - covers the binding site

Actin - slides toward the M line during a contraction

Troponin - binds Ca++ & starts the contraction cycle

Types of Skeletal Muscle Fibers:

Oxidative fibers—use aerobic pathways

Glycolytic fibers—use anaerobic glycolysis

Slow Oxidative Fibers - most resistant to fatigue

Fast Glycolytic Fibers - have few mitochondria

Fast Oxidative Fibers - contract quickly & rely on aerobic respiration for ATP

Connective Tissue Sheaths of Skeletal Muscles:

Perimysium - surrounds each muscle bundle (fascicle)

Epimysium - surrounds entire muscle (dense layer of collagen fibers)

Endomysium - surrounds each muscle fiber

Deep fascia - binds muscles into functional groups

Cellular Components of Skeletal Muscle Fibers:

Myofibril - rod-like contractile elements within a muscle fiber 80% of cell volume, they

have striations: aligned repeating series of dark A bands and light I bands

Sarcomere - smallest functional unit of muscle fiber, region of myofibril between Z discs,

composed of thick and thin myofilaments

Sarcoplasmic Reticulum - series of membranous channels (modified ER) that

surround each myofibril

Sarcoplasm - cytoplasm of a skeletal muscle fiber

Sarcolemma - plasma membrane of a skeletal muscle fiber

Skeletal Muscle Fiber Areas:

H Band - area in the center of the A band containing only thick filaments

A Band - area of the sarcomere with overlapping thick and thin filaments

I Band - area of the sarcomere containing only thin filaments

Z disc - sheets of proteins that anchor thin filaments & connect myofibrils

H zone - lighter mid-region, no overlapping filaments

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M line - line of protein myomesin that holds adjacent thick filaments together

Muscle Actions:

Wave Summation - when a muscle is stimulated repeatedly for several seconds with a

constant stimulus, the amount of tensions gradually increases to maximum.

Twitch - type of contraction represented by a single stimulus/relaxation sequence

Complete Tetanus - a muscle that is stimulated so frequently that the relaxation phase is

completely eliminated

Incomplete Tetanus - a muscle producing peak tensions with visible relaxation during

rapid cycles of contraction and relaxation.

Activities for Muscle Actions:

Endurance-type activities - best suited for slow oxidative fibers

Short-term intense movements - best for fast glycolytic fibers

400M or 800M sprint - best for fast oxidative fibers

Imbalances:

Strain - excessive stretching & possible tearing caused by overuse or abuse

Cramp - sustained spasm or tetanic contraction

Myalgia - pain resulting from any muscle disorder

Fibromyositis - inflammation of muscle, connective tissue coverings, tendons, and

capsules of nearby joints

Muscle Twitch Periods:

Latent Period - time b/n the stimulus/electrical event & the mechanical event of

contraction

Contraction Period - time during which the muscle is shortening

Refractory Period - very brief time after one stimulus during which the muscle is

unresponsive to a second stimulus

Relaxation Period - time during which the muscle is returning to its original length

Recovery Period - the body's need for oxygen is increased

Action Potential Information:

Resting Potential - initial polarized state

Action Potential - propagation of an electrical current along the sarcolemma

End Plate Potential - electrical event occurring only at neuromuscular junctions

Refractory Period - time when a fiber cannot be stimulated until repolarization is done

Repolarization - restoration of membrane potential to resting potential

Generation & Propagation of an Action Potential:

*Resting Sarcolemma Electrical Conditions are positive outside, negative inside

*Production of an end plate potential and depolarization of adjacent areas is the

depolarization & generation of action potential

*Increased positive charge inside sarcolemma changes permeability of adjacent areas,

opening voltage-regulated Na+ channels which is the propagation of the action potential

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*A change in sarcolemma after the wave of depolarization; Na+ channels close and K_

channels open allowing K+_ to create a positive charge outside the membrane is

repolarization

Force of Muscle Contractions Affected By:

Number of fibers

Relative size of the fibers—hypertrophy of cells increases strength

Frequency of stimulation— frequency allows time for more effective transfer of tension

to noncontractile components

Length-tension relationship—muscles contract most strongly when muscle fibers are 80–

120% of their normal resting length

 load   latent period,  contraction, and  duration of contraction

Developmental Aspects:

All muscle tissues develop from embryonic myoblasts

Myoblast-like skeletal muscle satellite cells have limited regenerative ability

Injured heart muscle is mostly replaced by connective tissue

Smooth muscle regenerates throughout life

Female skeletal muscle makes up 36% of body mass

Male skeletal muscle makes up 42% of body mass, primarily due to testosterone

Body strength per unit muscle mass is the same in both sexes

Terms to Know:

Synaptic Cleft - Space between the neuron and the muscle

Myosin molecules - cross bridges

Synaptic Knob - contains vesicles filled with acetylcholine

Motor End plate - contains receptors for acetylcholine

Axon Terminals forms neuromuscular junctions with a single muscle fiber

Motor Unit - a motor neuron & all the fibers it supplies

Cross Bridges - a link between the thick and thin filaments formed by the globular head

of thick filaments.

ATP - provides 95% of energy needed for contraction during moderate exercise

Myasthenia Gravis - autoimmune disease of acetylcholine receptors

Peristalsis - alternating contraction and relaxation of opposing layers of smooth muscle

Excitation-Contraction Coupling - events that form the link between electrical activity in

the sarcolemma and the initiation of a contraction

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