Superfamily SCARABAEOIDEA Latreille, 1802

PASSALID LINEAGE

GLAPHYRID LINE

Family LUCANIDAE Latreille, 1804

Subfamily LUCANINAE Latreille, 1804

Tribe LUCANINI Latreille, 1804

Genus Eurytrachelus Motschulsky, 1850. E. rectus (1) (AFROTROPICAL) – 1

Genus Lucanus Scopoli, 1763. L. capreolus (Linnaeus) (4), elaphus (Fabricius) (6), mazama (LeConte) (8), placidus (Say) (5) (NEARCTIC) – 23

Undet. sp. ex. Korea (1) (PALEARCTIC) – 1

Subfamily DORCINAE Parry, 1864

Tribe DORCINI Parry, 1864

Genus Dorcus MacLeay, 1819. D. parallelus (Say) (7) (NEARCTIC) – 7

Subfamily PLATYCERINAE Mulsant, 1842

Tribe PLATYCERINI Mulsant, 1842

Genus Platycerus Geoffrey, 1762. P. virescens (Fabricius) (6) (NEARCTIC); acuticollis Kurosawa (2), sp. ex. Korea (1) (PALEARCTIC) – 9

Subfamily NICAGINAE LeConte, 1861

Tribe NICAGINI LeConte, 1861

Genus Nicagus LeConte, 1861. N. obscurus (LeConte) (1) (NEARCTIC) – 1

Subfamily SYNDESINAE MacLeay, 1819

Tribe CERUCHINI Jacquelin du Val & Fairmaire, 1859

Genus Ceruchus MacLeay, 1819. C. piceus (Weber) (5) (NEARCTIC) – 5

Subfamily CHIASOGNATHINAE Burmeister, 1847

Tribe CHIASOGNATHINI Burmeister, 1847

Genus Chiasognathus Stephens, 1831. C. granti Stephens (1) (NEOTROPICAL) – 1

Family PASSALIDAE Leach, 1815

Subfamily PASSALINAE Leach, 1815

Tribe PASSALINI Leach, 1815

Genus Odontotaenius Kuwert, 1896. O. disjunctus (Illiger) (6) (NEARCTIC) – 6

Undet. spp. ex. Argentina, Brazil, Ecuador (15) (NEOTROPICAL) – 15

Family TROGIDAE MacLeay, 1819

Subfamily TROGINAE MacLeay, 1819

Genus Trox Fabricius, 1775. T. aequalis Say (4), affinis Robinson (1), hamatus Robinson (18), plicatus Robinson (10), robinsoni Vaurie (5), sonorae LeConte (3), spinulosus spinulosus Robinson (10), spinulosus dentibius Robinson (8), tuberculatus (DeGeer) (4), unistriatus Palisot de Beauvois (8), variolatus Melsheimer (1), sp. ex Florida (4) (NEARCTIC) – 75

Subfamily OMORGINAE Nikolajev, 2005

Genus Omorgus Erichson, 1847

Subgenus Omorgus (s. str.). O. asper LeConte (14), carinatus (Loomis) (4), inflatus (Loomis) (3), monachus (Herbst) (8), punctatus (Germar) (6), suberosus (Fabricius) (13) (NEARCTIC) – 48

Subgenus Omorgus (Afromorgus) Scholtz, 1986. O. sp. nr. gemmatus (Olivier) (1); sp. nr. squalidus (Olivier) (1) (AFROTROPICAL) – 2

Family PLEOCOMIDAE LeConte, 1861

Subfamily PLEOCOMINAE LeConte, 1861

Genus Pleocoma LeConte, 1856. P. fimbriata LeConte (1), tularensis Leach (3) (NEARCTIC) – 4

GEOTRUPID LINE

Family GEOTRUPIDAE Latreille, 1802

Subfamily BOLBOCERATINAE Mulsant, 1842

Tribe ATHYREINI Howden and Martínez, 1963

Genus Neoathyreus Howden and Martínez, 1963. N. sp. ex Argentina (1) (NEOTROPICAL) – 1

Tribe BOLBOCERATINI Mulsant, 1842

Genus Bolbapium Boucomont, 1911. B. sp. ex Brazil (1) (NEOTROPICAL) – 1

Genus Bolboceras Kirby, 1819. B.thoracicornis (Wallis) (1) (NEARCTIC) – 1

Genus Bolbocerastes Cartwright, 1953. B. imperialis imperialis Cartwright (2) (NEARCTIC) – 2

Genus Bolbocerosoma Schaeffer, 1906. B. biplagiatum Dawson & McCollock (1), farctum (Fabricius) (2), hamatum Brown (1), lepidissimum Brown (1) (NEARCTIC) – 5

Genus Odontaeus Samouelle, 1819. O. filicornis (Say) (2), obesus LeConte (1) (NEARCTIC) – 3

Tribe EUCANTHINI Nikolajev, 2002

Genus Eucanthus Westwood, 1848. E. lazarus (Fabricius) (5), subtropicus Howden (1) (NEARCTIC) – 6

Subfamily GEOTRUPINAE Latreille, 1802

Tribe GEOTRUPINI Latreille, 1802

Genus Geotrupes Latreille, 1796

Subgenus Geotrupes (Cnemotrupes) Jekel, 1865. G. blackburnii excrementi Say (24), splendidus splendidus (Fabricius) (1), splendidus miarophagus Say (18) (NEARCTIC) – 43

Subgenus Geotrupes (Geohowdenius) Zunino, 1984. G. opacus Haldeman (2) (NEARCTIC) – 2

Subgenus Geotrupes (Anoplotrupes) Jeckel, 1865. G. stercorosus (Hartmann, in Scriba) (1) (PALEARCTIC) – 1

Genus Peltotrupes Blanchard, 1888. P. profundus (Howden) (2) (NEARCTIC) – 2

Tribe LETHRINI Mulsant & Rey, 1871

Genus Lethrus Scopoli, 1777. L. apterus (2) (PALEARCTIC) – 2

Family OCHODAEIDAE Mulsant & Rey, 1871

Subfamily OCHODAEINAE Mulsant & Rey, 1871

Tribe OCHODAEINI Mulsant & Rey, 1871

Genus Neochodaeus Nikolayev, 1995. N. frontalis (LeConte) or n. sp. near (1), praesidii (Bates) (1) (NEARCTIC) – 2

Genus Parochodaeus Nikolyev, 1995. P. biarmatus (LeConte) (7), duplex (LeConte) (3), pectoralis (LeConte) (3) (NEARCTIC) – 13

Genus Xenochodaeus Paulsen, 2007. X. americanus (Westwood) (1), musculus (Say) (1), planifrons (Schaeffer) (4) (NEARCTIC) – 6

Undet. spp. ex Kansas, Missouri (3) (NEARCTIC) – 3

Family HYBOSORIDAE Erichson, 1847

Subfamily HYBOSORINAE Erichson, 1847

Genus Hybosorus MacLeay, 1847. H. illigeri Reiche (4) (NEARCTIC); illigeri Reiche (2) (AFROTROPICAL) – 6

Genus Phaeochrous Laporte, 1840. P. madagascariensis (Westwood) (7) (AFROTROPICAL) – 7

Subfamily CERATOCANTHINAE Martínez, 1968

Tribe CERATOCANTHINI Martínez, 1968

Genus Gemarostes Paulian, 1982. G. aphodioides (Illiger) (5), globosus (Say) (10) (NEARCTIC) – 15

Undet. sp. ex Ecuador (1) (NEOTROPICAL) – 1

Family GLAPHYRIDAE MacLeay, 1819

Subfamily GLAPHYRINAE MacLeay, 1819

Genus Lichnanthe Burmeister, 1844. L. sp. ex California (1) (NEARCTIC) – 1

SCARABAEID LINEAGE

Family SCARABAEIDAE Latreille, 1802

Subfamily APHODIINAE Leach, 1815

Tribe APHODIINI Leach, 1815

Subtribe APHODIINA Leach, 1815

Genus Aphodius Illiger, 1798

Subgenus Aphodius (s. str.). A. fimetarius (Linnaeus) (7) (NEARCTIC) – 7

Subgenus Aphodius (Labarrus) Mulsant and Rey, 1870. A. lividus (Olivier) (1) (NEARCTIC) – 1

Subgenus Aphodius (Colobopterus) Mulsant, 1842. A. erraticus (Linnaeus) (7) (NEARCTIC) – 7

Subfamily SCARABAEINAE Latreille, 1802

Tribe GYMNOPLEURINI Lacordaire, 1856

Genus Allogymnopleurus Janssens, 1940. A. thalassinus (Klug) (17) (AFROTROPICAL) – 17

Genus Garreta Janssens, 1940. G. nitens (Olivier (3) (AFROTROPICAL) – 3

Genus Gymnopleurus Illiger, 1803

Subgenus Gymnopleurus (s. str.). G. aenescens Wiedemann (1), virens Erichson (10) (AFROTROPICAL) – 11

Tribe CANTHONINI van Lansberge, 1874

Genus Anachalcos Hope, 1837. A. convexus Boheman (3) (AFROTROPICAL) – 3

Genus Canthon Hoffmanseg, 1817

Subgenus Canthon (s. str.). C. chalcites Haldeman (12), cyanellus LeConte (23), imitator Brown (10), indigaceus indigaceus LeConte (5), pilularius (Linnaeus) (34) (NEARCTIC); aberrans Harold (9), humectus incisus Robinson (4), humectus sayi Robinson (4), indigaceus chevrolati Harold (8), ?virens (Mannerheim) (2), sp. ex México [humectus-species group] (2), sp. ex Ecuador (2) (NEOTROPICAL) – 116

Subgenus Canthon (Boreocanthon) Halffter, 1958. C. lecontei Harold (9), simplex (LeConte) (1) (NEARCTIC); ?anteuchiceps Bates (1) (NEOTROPICAL) – 11

Subgenus Canthon (Glaphyrocanthon) Martínez, 1948. C. viridis (Palisot de Beauvois) (7) (NEARCTIC); ?leechi (Martinez, Halffter & Halffter) (3), ?luteicollis [sic for luteicornis?] Erichson (1), sp. ex México [viridis complex ♀] (1) (NEOTROPICAL) – 12

Genus Deltochilum Eschscholtz, 1822

Subgenus Deltochilum (Hybomidium) Shipp, 1897. D. gibbosum gibbosum (Fabricius) (5) (NEARCTIC) – 5

Genus Melanocanthon Halffter, 1958. M. nigricornis (Say) (27), punctaticollis (Schaeffer) (2), sp. nr. bispinatus Robinson ex Nebraska (1), sp. ex. Missouri (1) (NEARCTIC) – 31

Genus Pseudocanthon Bates, 1887. P. perplexus (LeConte) (14) (NEARCTIC) – 14

Tribe COPRINI Leach, 1815

Genus Catharsius Hope, 1837. C. sp. ex South Africa (2) (AFROTROPICAL) – 2

Genus Copris Geoffrey, 1762. C.arizonensis Schaeffer (2), fricator fricator (Fabricius) [= tullius Olivier] (10), inemarginatus Blatchley (2), lugubris Boheman (2), remotus remotus LeConte (5) (NEARCTIC); lugubris Boheman (1) (NEOTROPICAL); elphenor Klug (3), sp. ex South Africa (2) (AFROTROPICAL) – 27

Genus Dichotomius Hope, 1838

Subgenus Dichotomius (s. str.). D. colonicus (Say) (2), carolinus (Linnaeus) (5) (NEARCTIC); annae Kohlmann & Solis (1), colonicus (Say) (1), sp. ex Brazil (9), sp. ex Ecuador (1) (NEOTROPICAL) – 20

Genus Heliocopris Hope, 1837. H. sp. ex. India (1) (INDOMALAYAN) – 1

Genus Metacatharsius Paulian, 1939. M. spp. ex South Africa (2) (AFROTROPICAL) – 2

Genus Ontherus Erichson, 1847

Subgenus Ontherus (s. str.). O. sulcator (Fabricius) (1) (NEOTROPICAL) – 1

Tribe ATEUCHINI Laporte, 1840

Genus Ateuchus Weber, 1801. A. histeroides Weber (30), texanus Robinson (6) (NEARCTIC); rodriguezi (Borre) (4) (NEOTROPICAL) – 45

Genus Pedaria Laporte, 1832. P. sp. ex South Africa (4) (AFROTROPICAL) – 4

Genus Uroxys Westwood, 1842. U. sp. ex Ecuador (5) (NEOTROPICAL) – 5

Tribe ONITICELLINI d'Orbigny, 1916

Subtribe ONITICELLINA Kolbe, 1905

Genus Euoniticellus Janssens, 1953. E. intermedius (Reiche) (3) (NEARCTIC); intermedius (Reiche) (1) (AFROTROPICAL) – 1

Tribe ONITINI Laporte, 1840

Genus Onitis Fabricius, 1798. O. fulgidus Klug (3), inversidens Lansberge (1), pseudosetosus Ferreira (1), uncinatus Klug (5) (AFROTROPICAL) – 10

Tribe ONTHOPHAGINI Burmeister, 1846

Genus Caccobius Thomson, 1859.

Subgenus Caccobius (Caccophilus) Jekel, 1872. C. ferrugineus (Fahraeus) (46) (AFROTROPICAL) – 46

Subgenus ?? C. sp. ex South Africa (4) (AFROTROPICAL) – 4

Genus Cleptocaccobius Cambefort, 1984. C. sp. ex South Africa (1) (AFROTROPICAL) – 1

Genus Digitonthophagus Balthasar, 1959. D. gazella (Fabricius) (21) (NEARCTIC); gazella (Fabricius) (9) (AFROTROPICAL) – 30

Genus Onthophagus Latreille, 1802

Subgenus Onthophagus (s. str.). O. aciculatus Blatchley (5), cavernicollis Howden & Cartwright (1-1 deposited WBWC), depressus Harold (2), hecate hecate (Panzer) (58), hecate blatchleyi Brown (3), hopfneri Harold (3), nuchicornis (Linnaeus) (3), orpheus pseudorphaeus Howden & Cartwright (8), pennsylvanicus Harold (40), schaefferi Howden & Cartwright (1), striatulus striatulus (Palisot de Beauvois) (4), subaeneus (Palisot de Beauvois) (7), taurus (Schreber) (83), velutinus Horn (5) (NEARCTIC); ?batesi Howden & Cartwright (1), hopfneri Harold (4), ?incensus Say (1), sp. ex México [landolti-species group] (2), sp. ex. Mexico (6) (NEOTROPICAL); apiciosus d’Orbigny (3), bicavifrons d’Orbigny (4), carbonarius Klug (6), flavolimbatus Klug (2), fimetarius Roth (5), ochropygus d’Orbigny (5), pallidipennis Fåhraeus (2), quadrinodosus Fåhraeus (1), signatus Fåhraeus (5), variegatus (Fabricius) (4), vinctus Erichson (30), sp. ex South Africa (1) (AFROTROPICAL) – 304

Genus Phalops Erichson, 1848. P. ardea Klug (1), boschas Klug (2) (AFROTROPICAL) – 3

Genus Proagoderus Lansberge, 1883. P. sapphirinus (d’Orbigny, 1916) (1) (AFROTROPICAL) – 1

Tribe SCARABAEINI Latreille, 1802

Genus Kheper Janssens, 1940. K. clericus (Boheman) (10) (AFROTROPICAL) – 10

Genus Pachylomerus Bertoloni, 1849. P. femoralis Kirby (11) (AFROTROPICAL) – 11

Genus Scarabaeus Linnaeus, 1758. S. galenus (Westwood) (6), goryi (Laporte) (1), sp. ex South Africa (1) (AFROTROPICAL) – 8

Tribe PHANAEINI Kolbe, 1905

Genus Coprophanaeus d’Ousoufieff, 1924

Subgenus Coprophanaeus (s. str.). C. milon milon (Blanchard) (1) (NEOTROPICAL) – 1

Genus Phanaeus MacLeay, 1819

Subgenus Phanaeus (s. str.). P. quadridens Say (5), triangularis triangularis Say (1), vindex MacLeay (6) (NEARCTIC); daphnis Harold (4), sp. ex México [poss. demon Laporte] (1) (NEOTROPICAL) – 17

Genus Oxysternon Laporte, 1840. O. conspicilatum (Weber) (1) (NEOTROPICAL) – 1

Genus Sulcophanaeus d’Ousoufieff, 1924. S. menelas (Laporte) (1) (NEOTROPICAL) – 1

Tribe SISYPHINI Mulsant, 1842

Genus Sisyphus Latreille, 1807. S. costatus (Thunberg) (13), impressipennis Lansberge (2) (AFROTROPICAL) – 15

Subfamily MELOLONTHINAE Leach, 1819

Tribe SERICINI Kirby, 1837

Subtribe SERICINA Kirby, 1837

Genus Ablaberoides Blanchard, 1850. A. sp. ex South Africa (1) (AFROTROPICAL) – 1

Genus Phylloserica Brenske, 1899. P. vittata Blanchard (1) (AFROTROPICAL) – 1

Genus Serica MacLeay, 1819. S. evidens Blatchley (3), mystaca Dawson (12), sp. ex Missouri (2) (NEARCTIC) – 17

Tribe DIPLOTAXINI Kirby, 1837

Genus Diplotaxis Kirby, 1837. D. frondicola (Say) (24), misella Fall (6), thoracica Fall (21), truncatula LeConte (4) (NEARCTIC) – 55

Tribe MELOLONTHINI Leach, 1819

Subtribe MELOLONTHINA Leach, 1819

Genus Phyllophaga LeConte, 1856

Subgenus Phyllophaga (s. str.). P. balia (Say) (6), crenulata (Frölich) (3), crinita (Burmeister) (7), futilis (LeConte) (1), hirticula (Knoch) (2), inversa (Horn) (6), quercus (Knoch) (2), ?rubiginosa (LeConte) (8), spreta (Horn) (3), subtonsa (LeConte) (1), sp. ex Missouri [tristis-species group] (1), spp. ex Texas (3) (NEARCTIC); forcipata (Burmeister) or near (1) (NEOTROPICAL) – 44

Subgenus Phyllophaga (Eugastra) LeConte, 1856. P. cribrosa (LeConte) (2) (NEARCTIC) – 2

Genus Polyphylla Harris, 1841

Subgenus Polyphylla (s. str.). P. gracilis Horn (2), occidentalis (Linnaeus) (1), pottsorum Hardy & Andrews (2), spp. ex. Arizona, California, Texas (13) (NEARCTIC) – 18

Tribe MACRODACTYLINI Kirby, 1837

Genus Dichelonyx Harris, 1827. D. albicollis (Burmeister) (6), backii (Kirby) [= fulgida LeConte] (1), diluta (Fall) (5), elongatula (Fabricius) (11), pusilla (LeConte) (3), subvittata (LeConte) (5), truncata (LeConte) (1), vandykei Saylor (2), vicina (Fall) (1), sp. ex Missouri (10) (NEARCTIC) – 45

Genus Isonychus Mannerheim, 1829. I. arizonensis Howden (5) (NEARCTIC) – 5

Genus Macrodactylus Dejean, 1821. M. angustatus (Palisot de Beauvois) (5), subspinosus (Fabricius) (5), uniformis Horn (13) (NEARCTIC); sp. ex Ecuador (1) (NEOTROPICAL) – 24

Tribe HOPLIINI Latreille, 1829

Subtribe HOPLIINA Latreille, 1829

Genus Hoplia Illiger, 1803

Subgenus Hoplia (s. str.). H. callipyge LeConte [= oregona LeConte] (10), dispar LeConte (10), modesta LeConte (5), trifasciata Say (5) (NEARCTIC) – 30

Subtribe ??

Undet. sp. ex South Africa (2) (AFROTROPICAL) – 2

Tribe ??

Undet. sp. ex South Korea (PALEARCTIC) – 4

Subfamily RUTELINAE MacLeay, 1819

Tribe RUTELINI MacLeay, 1819

Subtribe RUTELINA MacLeay, 1819

Genus Antichira Eschscholtz, 1818

Subgenus Antichira (Dorysthetus) Blanchard, 1845. A. fulgida Waterhouse (1) (NEOTROPICAL) – 1

Genus Calomacraspis Bates, 1888. C. splendens (Burmeister) (5) (NEOTROPICAL) – 5

Genus Chrysina Kirby, 1828. C. beyersi (Skinner) (14), gloriosa (LeConte) (23), lecontei (Horn) (6), woodi (Horn) (1) (NEARCTIC); strasseni (Ohaus) (1) (NEOTROPICAL) – 45

Genus Chrysophora Serville, 1825. C. chrysochloa (2) (NEOTROPICAL) – 2

Genus Cotalpa Burmeister, 1844. C. consobrina Horn (5), lanigera (Linnaeus) (5) (NEARCTIC) – 10

Genus Macraspis MacLeay, 1819. M. pantochloris Blanchard (4) (NEOTROPICAL) – 4

Genus Paracotalpa Ohaus, 1915. P. granicollis (Haldeman) (1), puncticollis (LeConte) (1), ursina (Horn) (102) (NEARCTIC) – 104

Genus Pelidnota MacLeay, 1819. P. lugubris LeConte (5), punctata (Linnaeus) (6) (NEARCTIC); virescens Burmeister (6) (NEOTROPICAL) – 17

Genus Rutela Latreille, 1802. R. lineola (Linnaeus) (6) (NEOTROPICAL) – 6

Tribe ANOMALINI Streubel, 1839

Subtribe ANOMALINA Streubel, 1839

Genus Anomala Samouelle, 1819. A. binotata (Gyllenhal) (6), delicata Casey (10), flavipennis flavipennis Burmeister (3), innuba (Fabricius) (16), insitiva Robinson (3), lucicola (Fabricius) (7), marginata (Fabricius) (5), nimbosa Casey (1), sabinae Potts (5), undulata Melsheimer (9) (NEARCTIC); histrionella Bates (4), testaceipennis Blanchard (1), villosela Blanchard (2) (NEOTROPICAL); sp. ex South Korea (1) (PALEARCTIC); expansa Bates (1) (INDOMALAYAN) – 74

Genus Mimela Kirby, 1825. M. taiwana Sawada (2) (INDOMALAYAN) – 2

Genus Phyllopertha Stephens, 1830. P. diversa Waterhouse (2) (PALEARCTIC) – 2

Genus Strigoderma Burmeister, 1844. S. arbicola (Fabricius) (13), pimalis Casey (4), pygmaea (Fabricius) (5) (NEARCTIC); costulipennis Bates (2), sulcipennis Burmeister (3), vestita Burmeister (2), villosella Blanchard (4) (NEOTROPICAL) – 33

Subtribe POPILIINA Ohaus, 1918

Genus Peripopillia Kolbe, 1894. P. basalis (Blanchard) (1) (AFROTROPICAL) – 1

Genus Popillia Dejean, 1821. P. japonica Newman (7) (NEARCTIC); bipunctata (Fabricius) (1) (AFROTROPICAL) – 8

Subtribe ANISOPLIINA Burmeister, 1844

Genus Anomalacra Casey, 1915. A. clypealis (Schaeffer) (2) (NEARCTIC) – 2

Tribe ADORETINI Burmeister, 1844

Subtribe ??

Undet. sp. ex South Africa (1) (AFROTROPICAL) – 1

Subfamily DYNASTINAE MacLeay, 1819

Tribe CYCLOCEPHALINI Laporte, 1840

Genus Ancognatha Erichson, 1847. A. manca (LeConte) (5) (NEARCTIC); vulgaris Arrow (5) (NEOTROPICAL) – 10

Genus Chalepides Casey, 1915. C. hydrophiloides (Burmeister) (1) (NEOTROPICAL) – 1

Genus Dyscinetus Harold, 1869. D. morator (Fabricius) (5) (NEARCTIC); dubius (Olivier) (1), sp. ex Argentina ♀♀ (3) (NEOTROPICAL) – 9

Tribe DYNASTINI MacLeay, 1819

Genus Allomyrina Arrow, 1911. A. dichotomus (Linnaeus) (1) (INDOMALAYAN) – 1

Genus Dynastes MacLeay, 1819. D. granti Horn (1), tityus (Linnaeus) (12) (NEARCTIC); hercules (Linnaeus) (7), neptunus Quenzel (1) (NEOTROPICAL) – 21

Genus Golofa Hope, 1837. G. unicolor (Bates) (1), eacus Burmeister (1) (NEOTROPICAL) – 2

Genus Megasoma Kirby, 1825. M. actaeon actaeon (Linnaeus) (37) (NEOTROPICAL) – 37

Tribe ORYCTINI Mulsant, 1842

Genus Coelosis Kirby & Spence 1828. C. biloba (Linnaeus) (1) (NEOTROPICAL) – 1

Genus Enema Hope, 1837. E. endymion Chevrolat (4), pan (Fabricius) (6) (NEOTROPICAL) – 10

Genus Heterogomphus Burmeister, 1847. H. bourcieri Guérin (1), dilaticollis Burmeister (1), sp. ex. Ecuador (1) (NEOTROPICAL) – 3

Genus Megaceras Hope, 1837. M. sp. ex Ecuador (2) (NEOTROPICAL) – 2

Genus Oryctes Illiger, 1798. O. rhinocerus (1) (INDOMALAYAN) – 1

Genus Strategus Kirby, 1828. S. aloeus julianus Burmeister (10), antaeus (Drury) (2), cessus LeConte (6) (NEARCTIC); surinamensis surinamensis (Burmeister) (5), surinamensis hirtus Sternberg (1) (NEOTROPICAL) – 23

Genus Xyloryctes Hope, 1837. X. jamaicensis (Drury) (8) (NEARCTIC) – 8

Tribe PENTODONTINI Mulsant, 1842

Subtribe PENTODONTINA Mulsant, 1842

Genus Bothynus Hope, 1837. B. cunctator (Mannerheim) (11), medon (Germar) (3), striatellus (Fairmaire) (1) (NEOTROPICAL) – 15

Genus Diloboderus Reiche, 1859. D. abderus (Sturm) (15) (NEOTROPICAL) – 15

Genus Eutheola Bates, 1888. E. rugiceps (LeConte) (11) (NEARCTIC) – 11

Genus Temnorhynchus Hope 1837

Subgenus Temnorhynchus (s. str.). T. coronatus (Fabricius) (7) (AFROTROPICAL) – 7

Genus Tomarus Erichson, 1847. T. gibbosus (DeGeer) (7), relictus (Say) (5) (NEARCTIC); bituberculatus (Beauvois) (1), gyas Erichson (2) (NEOTROPICAL) – 15

Tribe PHILEURINI Burmeister, 1847

Subtribe PHILEURINA Burmeister, 1847

Genus Archophileurus Kolbe, 1910. A. sp. ex Argentina (2) (NEOTROPICAL) – 2

Genus Hemiphileurus Kolbe, 1910. H. illatus illatus (LeConte) (7) (NEARCTIC); illatus mexicanus Endrödi (1) (NEOTROPICAL) – 8

Genus Homophileurus Kolbe 1910. H. quadrituberculatus (Palisot de Beauvois) (2) (NEOTROPICAL) – 2

Genus Phileurus Latreille, 1807. P. trunctatus (Palisot de Beauvois) (5), valgus (Linnaeus) [= castaneus Haldeman] (20) (NEARCTIC); carinatus Prell (5) (NEOTROPICAL) – 14

Subfamily CETONIINAE Leach, 1815

Tribe VALGINI Mulsant, 1842

Subtribe VALGINA Mulsant, 1842

Genus Valgus Scriba, 1790. V. canaliculatus (Olivier) (15) (NEARCTIC) – 15

Tribe TRICHIINI Fleming, 1821

Subtribe TRICHIINA Fleming, 1821

Genus Apeltastes Howden, 1968. A. elongata Howden (2) (NEOTROPICAL) – 2

Genus Gnorimella Casey, 1915. G. maculosa (Knoch) (2) (NEARCTIC) – 2

Genus Myodermum Burmeister and Schaum, 1840. M. rufum Waterhouse (2) (AFROTROPICAL) – 2

Genus Trichiotinus Casey, 1915. T. affinis (Gory & Percheron) (3), assimilis (Kirby) (5), bibens (Fabricius) (5), lunulatus (Fabricius) (15), piger (Fabricius) (16), rufobrunneus (Casey) (7), texanus (Horn) (9), viridans (Kirby) (12) (NEARCTIC) – 72

Genus Trichius Fabricius, 1775. T. succinctus (Pallas) (1) (PALEARCTIC) – 1

Genus Trigonopeltastes Burmeister, 1840. T. delta (Forster) (19) (NEARCTIC), archmedes Shaum (11), geometrica Schaum (15) (NEOTROPICAL) – 45

Subtribe OSMODERMATINA Schenkling, 1922

Genus Osmoderma LePeletier & Serville, 1828. O. subplanata Casey (4) (NEARCTIC) – 4

Tribe CREMASTOCHEILINI Burmeister & Schaum, 1841

Subtribe OPLOSTOMATINA Krikken, 1984

Genus Oplostomus MacLeay,1838. O. fuligineus (Olivier) (6) (AFROTROPICAL) – 6

Subtribe CREMASTOCHEILINA Burmeister & Schaum, 1841

Genus Campsiura Hope, 1831. O. sp. ex South Africa (4, on loan to Bruce Gill) (AFROTROPICAL) – 4

Genus Cremastocheilus Knoch, 1801

Subgenus Cremastocheilus (s. str.). C. castaneus castaneus Knoch (7), nitens LeConte (1) (NEARCTIC) – 8

Subgenus Cremastocheilus (Myrmecotonus) Mann, 1914. C. angularis LeConte (1) (NEARCTIC) – 1

Subgenus Cremastocheilus (??). C. sp. ex California (1), sp. ex Missouri (7) (NEARCTIC) – 8

Tribe GOLATHINI Griffith & Pidgeon, 1832

Subtribe CORYPHOCERINA Burmeister, 1842

Genus Anisorrhina Westwood, 1842.

Subgenus Anisorrhina (Melinesthes) Kraatz, 1880. A. algoensis (Westwood) (12) (AFROTROPICAL) – 12

Genus Eudicella White, 1839. E. gralli (Buquet) (2), smithii (MacLeay) (9) (AFROTROPICAL) – 11

Genus Plaesiorrhynella Krikken, 1984.

Subgenus Plaesiorrhynella (Chondrorrhina) Kraatz, 1880. P. trivittata (Schaum) (18) (AFROTROPICAL) – 18